Systématique / Les Lucanidae Du Monde / Typus Materials / Typus Bartolozzi Luca Collection / Sphaenognathus Genera / SPHAENOGNATHUS MATHEUSI ( PARATYPUS )

Systématique [14859]
- Les Lucanidae Du Monde [7253]
- Les Dynastidae Du Monde [5287]
  - Introduction [1]
  - Dynastidae GENERA [5092]
    - AGAOCEPHALINI [137]
      - AEGOPSIS Burmeister, 1847 [13]
        bolboceridus (Thomson, 1860) [4]
        chaminadei Dechambre, 1999 [1]
        curvicornis Burmeister, 1847 [3]
        peruvianus Arrow, 1941 [1]
        westwoodi Thomson, 1860 [3]
      - AGAOCEPHALA Serville, 1825 [25]
        bicuspis Erichson, 1848 [3]
        brasiliana Martinez & Alvarenga, 1987 [1]
        cornigera Serville, 1828 [3]
        duponti Laporte de Castelnau, 1832 [3]
        inermicollis Arrow, 1914 [3]
        mannerheimi Laporte de Castelnau 1832 [3]
        margaridae Alvarenga, 1958 [4]
        melolonthida Thomson, 1860 [1]
        urus Thomson, 1860 [3]
      - ANTODON Brême, 1844 [3]
        goryi (Laporte de Castelnau, 1832) [3]
      - BRACHYSIDERUS Waterhouse, 1881 [22]
        elyanae Dechambre, 2009 [3]
        goyanus Ohaus, 1930 [1]
        martinae Abadie, 2010 [1]
        matogrossensis Ohaus, 1930 [3]
        mielkeorum Grossi & Grossi, 2005 [3]
        minicola Ohaus, 1930 [3]
        paranensis lenorae Grossi, Dechambre & Grossi, 2012 [1]
        paranensis paranensis Arrow, 1902 [1]
        quadrimaculatus quadrimaculatus Waterhouse, 1881 [4]
        quadrimaculatus tridentiger Prell, 1934 [1]
      - COLACUS Ohaus, 1910 [3]
        bicolor Ohaus, 1910 [1]
        endrödii Martinez, 1987 [1]
        morio Ohaus, 1910 [1]
      - DEMOCRATES Burmeister, 1847 [9]
        burmeisteri Reiche, 1852 [4]
        croesus (Newman, 1836) [3]
        lepiouffi Dechambre, 2006 [1]
      - GNATHOGOLOFA Arrow, 1914 [3]
        bicolor (Ohaus, 1910) [3]
      - HORRIDOCALIA Endrödi, 1974 [1]
        delislei Endrödi, 1974 [1]
      - LYCOMEDES Brême, 1844 [23]
        buckleyi Waterhouse, 1880 [4]
        burmeisteri Waterhouse, 1880 [1]
        hirtipes Arrow, 1902 [4]
        ohausi Arrow, 1908 [4]
        ramosus Arrow, 1902 [1]
        reichei Brême, 1844 [4]
        velutipes Arrow, 1902 [4]
      - MITRACEPHALA Thomson, 1859 [6]
        humboldti Thomson, 1859 [4]
        lachaumei Dechambre, 1992 [1]
      - SPODISTES Burmeister, 1847 [29]
        angulicollis Dechambre, 1992 [1]
        armstrongi Dechambre, 1994 [4]
        batesi Arrow, 1902 [4]
        beltianus Bates, 1887 [4]
        grandis Sternberg, 1903 [3]
        hopei Arrow, 1902 [4]
        mniszechi (Thomson, 1860) [4]
        monzoni Warner, 1992 [4]
    - CYCLOCEPHALINI [1137]
      - ACROBOLBIA Ohaus, 1912 [1]
        macrophylla Ohaus, 1912 [1]
      - ANCOGNATHA Erichson, 1847 [76]
        atacazo (Kirsch, 1885) [3]
        castanea Erichson, 1847 [3]
        erythrodera (Blanchard, 1846) [6]
        falsa Arrow, 1911 [4]
        gracilis Endrödi, 1966 [3]
        horrida Endrödi, 1967 [4]
        humeralis (Burmeister, 1847) [4]
        hyltonscottae Martinez, 1965 [3]
        jamesoni Murray, 1857 [6]
        lutea Erichson, 1847 [4]
        manca (LeConte, 1866) [4]
        matilei Dechambre, 2000 [1]
        nigriventris Otoya, 1945 [1]
        quadripunctata Bates, 1887 [4]
        rugulosa Endrödi, 1966 [1]
        scarabaeoides Erichson, 1847 [4]
        sellata Arrow, 1911 [3]
        ustulata (Burmeister, 1847) [7]
        veliae Pardo-Locarno-et-al, 2006 [1]
        vexans Ratcliffe, 1992 [4]
        vulgaris Arrow, 1911 [5]
      - ARRIGUTTIA Martinez, 1968 [5]
        bolivari Martinez, 1968 [1]
        brevissima (Arrow, 1911) [3]
      - ASPIDOLEA Bates, 1888 [69]
        bleuzeni Dechambre, 1992 [4]
        boulardi Dechambre, 1992 [1]
        brunnea Höhne, 1922 [1]
        chalumeaui Endrödi, 1977 [1]
        clypeata (Burmeister, 1847) [4]
        cognata Höhne, 1922 [3]
        collaris Endrödi, 1966 [1]
        ecuadoriana Endrödi, 1985 [4]
        epipleuralis Höhne, 1922 [1]
        fuliginea (Burmeister, 1847) [4]
        gaudairethorei Endrödi, 1980 [5]
        helleri (Höhne, 1922) [4]
        kuntzeni Höhne, 1922 [1]
        laticeps (Harold, 1869) [4]
        lindae Ratcliffe, 1977 [1]
        mimethes (Höhne, 1922) [1]
        notaticollis Höhne, 1922 [8]
        pelioptera (Burmeister, 1847) [4]
        quadrata Endrödi, 1980 [4]
        singularis Bates, 1887 [4]
        suturalis (Höhne, 1922) [2]
        suturella (Höhne, 1922) [1]
        testacea (Höhne, 1922) [3]
        theresae Dupuis, 2000 [1]
        vicina Dechambre, 1992 [1]
      - AUGODERIA Burmeister, 1847 [9]
        boliviana Endrödi, 1981 [1]
        freyi Endrödi, 1967 [1]
        ginglarisi Ponchel, 2009 [1]
        nitidula nitidula Burmeister, 1847 [4]
        nitidula yungana Martinez, 1966 [1]
      - CHALEPIDES Casey, 1915 [45]
        alliaceus (Burmeister, 1847) [3]
        anomalus Martinez, 1978 [1]
        barbatus argentinus Prell, 1938 [1]
        barbatus barbatus (Fabricius, 1781) [3]
        carinatus Joly & Escalona, 2002 [1]
        comes Prell, 1936 [4]
        dilatatus (Mannerheim, 1829) [3]
        dytiscoides (Arrow, 1911) [4]
        eucephalus (Casey, 1915) [1]
        fuliginosus (Burmeister, 1847) [6]
        howdenorum Joly & Escalona, 2002 [1]
        hydrophiloides (Burmeister, 1847) [4]
        luridus (Burmeister, 1847) [3]
        narcisoi Martinez, 1978 [1]
        osunai Joly & Escalona, 2002 [1]
        paradytis Ponchel & Dechambre 2003 [1]
        unicolor (Endrödi, 1963) [6]
      - CYCLOCEPHALA Dejean, 1821 [717]
        abrelata Ratcliffe & Cave, 2002 [1]
        acoma Ratcliffe, 2008 [1]
        aequatoria Endrödi, 1963 [3]
        affinis Endrödi, 1966 [1]
        alazonia Ratcliffe, 2003 [1]
        alexi Ratcliffe & Delgado, 1990 [1]
        almitana Dechambre, 1992 [4]
        altamontana Dechambre, 1999 [1]
        alutacea Höhne, 1923 [1]
        alvarengai Dechambre, 1980 [1]
        amazona amazona (Linnaeus, 1767) [4]
        amazona boliviensis Höhne, 1923 [1]
        amazona signata (Fabricius, 1781) [1]
        amblyopsis Bates, 1887 [7]
        ampliata Bates, 1887 [1]
        anibali Joly, 2009 [1]
        annamariae Dutrillaux & Chalumeau, 2013 [1]
        aravaipensis Ratcliffe, 1992 [1]
        arenosa Howden & Endrödi, 1966 [1]
        arnaudi Dechambre, 1980 [1]
        arrowiana Martinez, 1967 [1]
        atricapilla Mannerheim, 1829 [3]
        atriceps (Casey, 1915) [1]
        atricolor Chapin, 1932 [1]
        atripes (Bates, 1887) [1]
        barrerai Martinez, 1969 [1]
        batesi Delgado & Castaneda, 1994 [1]
        bella Endrödi, 1969 [1]
        berti Delgado, 1992 [1]
        bicolor Laporte de Castelnau, 1840 [4]
        bicolorata Endrödi, 1964 [1]
        bimaculata Dechambre, 1999 [1]
        binotata Dechambre, 1999 [1]
        bleuzeni Dechambre, 1995 [1]
        boliviana Dechambre, 1997 [1]
        bollei Dechambre & Endrödi, 1984 [1]
        borburatae Endrödi, 1980 [1]
        borealis Arrow, 1911 [3]
        boucheri Dechambre, 1997 [4]
        boulardi Dechambre, 1979 [1]
        brasiliana Endrödi, 1966 [1]
        brevipennis Endrödi, 1985 [1]
        brevis Höhne, 1923 [4]
        brittoni Endrödi, 1964 [3]
        burmeisteri Endrödi, 1964 [1]
        caelestis Ratcliffe & Delgado, 1990 [1]
        camachicola Ohaus, 1910 [6]
        capitata Höhne, 1923 [3]
        carbonaria Arrow, 1911 [1]
        cardini Chapin, 1935 [1]
        carinatipennis Martinez & M. 1984 [1]
        carlsoni Ratcliffe, 2008 [1]
        cartwighti Endrödi, 1964 [3]
        casanova Ratcliffe & Cave, 2009 [1]
        castanea (Olivier, 1789) [4]
        castaniella Bates, 1887 [1]
        caussaneli Dechambre, 1999 [4]
        cearae Höhne, 1923 [1]
        celata Dechambre, 1980 [1]
        chalumeaui Martinez, 1978 [1]
        chera Ratcliffe, 2008 [1]
        chiquita Ratcliffe, 2008 [1]
        colasi Endrödi, 1964 [4]
        collaris Burmeister, 1847 [3]
        comata Bates, 1887 [3]
        compacta Ratcliffe, 2008 [1]
        complanata Burmeister, 1847 [6]
        concolor Burmeister, 1847 [3]
        confusa Endrödi, 1966 [4]
        conspicua Sharp, 1877 [3]
        contraria Kirsch, 1873 [4]
        coriacea Dechambre, 1992 [1]
        couturieri Dechambre, 1999 [1]
        crassa Endrödi, 1967 [1]
        crepuscularis Martinez, 1954 [3]
        cribrata Burmeister, 1847 [4]
        cubana Chapin, 1932 [3]
        curta Bates, 1887 [3]
        dalensi Ponchel, 2009 [1]
        danforthi Chapin, 1935 [1]
        deceptor (Casey, 1915) [4]
        decorella Endrödi, 1966 [3]
        defecta Endrödi, 1968 [1]
        deltoides Ratcliffe, 1992 [1]
        dichroa Dechambre, 1992 [3]
        dilatata (Prell, 1934) [4]
        diluta Erichson, 1847 [4]
        discicollis Arrow, 1902 [1]
        discolor (Herbst, 1790) [6]
        dispar (Herbst, 1790) [3]
        distincta Burmeister, 1847 [3]
        divaricata Joly, 2005 [1]
        dolichotarsa Ratcliffe & Cave, 2008 [1]
        dominicana Endrödi, 1985 [1]
        duodecimpunctata Endrödi, 1966 [3]
        durantonorum Dechambre, 1999 [3]
        dyscinetoides Dechambre, 1999 [1]
        emarginata Endrödi, 1966 [3]
        endroedy-youngai Endrödi, 1964 [1]
        endrödii Martinez, 1965 [1]
        enigma Ratcliffe, 2003 [1]
        epistomalis Bates, 1887 [3]
        ergastuli Dechambre, 1997 [4]
        erotylina Arrow, 1914 [3]
        falsa Arrow, 1911 [1]
        fankhaeneli Endrödi, 1964 [3]
        fasciolata Bates, 1887 [3]
        ferruginea Fabricius, 1801 [1]
        figurata Burmeister, 1847 [1]
        flavipennis Arrow, 1914 [1]
        flavoscutellaris Höhne, 1923 [1]
        flora Arrow, 1911 [1]
        forcipulata Howden & Endrödi, 1966 [1]
        forsteri forsteri Endrödi, 1963 [1]
        forsteri maracayensis Endrödi, 1963 [1]
        forsteri mexicoi Endrödi, 1966 [1]
        freudei Endrödi, 1963 [3]
        freyi freyi Endrödi, 1964 [1]
        freyi integra Dechambre, 1999 [1]
        fulgurata Burmeister, 1847 [3]
        fulvipennis Burmeister, 1847 [4]
        fusiformis Chapin, 1932 [1]
        gabaldoni Martinez, 1981 [3]
        genieri Joly, 2010 [1]
        gigantea Dupuis, 2000 [1]
        goetzi Endrödi, 1966 [1]
        gravis Bates, 1887 [7]
        gregaria Heyne & Taschenberg, 1907 [4]
        guaguarum Dechambre & Endrödi, 1984 [4]
        guianae Endrödi, 1969 [4]
        guttata Bates, 1887 [4]
        guycolasi Dechambre, 1992 [1]
        halffteriana Martinez, 1968 [1]
        hardyi Endrödi, 1975 [3]
        hartmannorum Maly, 2006 [1]
        hayekae Endrödi, 1966 [1]
        helavai Endrödi, 1976 [1]
        herteli Endrödi, 1964 [1]
        hiekei Endrödi, 1964 [1]
        hielkemaorum Ratcliffe, 2008 [1]
        hirsuta Höhne, 1923 [1]
        hirta LeConte, 1861 [4]
        histrionica Burmeister, 1847 [3]
        holmbergei Martinez, 1968 [1]
        howdenannae Endrödi, 1976 [1]
        howdeni Endrödi, 1967 [1]
        husingi Endrödi, 1964 [1]
        iani Ratcliffe, 1992 [1]
        immaculata coahuilae Bates, 1887 [1]
        immaculata ferruginea (Fabricius, 1801) [1]
        immaculata immaculata (Olivier, 1789) [3]
        inca Endrödi, 1966 [1]
        insulicola Arrow, 1937 [1]
        isabellina Höhne, 1923 [4]
        isthmiensis Ratcliffe, 1992 [1]
        italoi Dupuis, 2000 [1]
        jalapensis Casey, 1915 [1]
        jaufferti Dechambre, 1979 [1]
        kahanoffae Martinez, 1975 [1]
        kaszabi Endrödi, 1964 [1]
        kechua (Martinez, 1957) [3]
        knobelae (Brown, 1934) [1]
        krombeini Endrödi, 1979 [1]
        kuntzeniana Höhne, 1923 [3]
        labidion Ratcliffe, 2003 [1]
        lachaumei Dechambre, 1992 [1]
        laevis Arrow, 1937 [1]
        lamarcki Dechambre, 1999 [1]
        laminata Burmeister, 1847 [4]
        larssoni Endrödi, 1964 [3]
        latericia Höhne, 1923 [4]
        latipennis Arrow, 1911 [1]
        latreillei Dechambre, 1999 [1]
        lecourti Dechambre, 1992 [1]
        letiranti Young, 1992 [1]
        lichyi Dechambre, 1980 [4]
        ligyrina Bates, 1887 [3]
        lineata Dupuis, 2008 [1]
        lineigera Höhne, 1923 [1]
        liomorpha Arrow, 1911 [1]
        literata Burmeister, 1847 [1]
        lizeri Martinez, 1964 [1]
        longa Endrödi, 1963 [1]
        longicollis Burmeister, 1847 [3]
        longimana Dechambre, 1980 [1]
        longitarsis Dechambre, 1999 [1]
        longula LeConte, 1863 [3]
        lunulata Burmeister, 1847 [4]
        lurida coahuilae Bates, 1888 [1]
        lurida lurida Bland, 1863 [3]
        lutea Endrödi, 1966 [4]
        macrophylla Erichson, 1847 [1]
        maculiventris Höhne, 1923 [1]
        maffafa Burmeister, 1847 [4]
        magdalenae Young & Le Tirant, 2005 [1]
        malleri Martinez, 1968 [1]
        mannheimsi Endrödi, 1964 [4]
        marginalis Kirsch, 1871 [4]
        marginicollis Arrow, 1902 [1]
        marianista Dechambre & Endrödi, 1984 [1]
        marqueti Dechambre, 1997 [1]
        martinezi Endrödi, 1964 [1]
        marylizae Ratcliffe, 2003 [1]
        mathani Dechambre, 1982 [1]
        mechae Martinez, 1978 [1]
        mecynotarsis Höhne, 1923 [4]
        megalophylla Endrödi, 1966 [1]
        meinanderi Endrödi, 1964 [3]
        melanae Bates, 1887 [1]
        melanocephala melanocephala (Fabricius, 1775) [3]
        melanocephala rubiginosa Burmeister, 1847 [2]
        melanopoda Ratcliffe, 2008 [1]
        melolonthida Ratcliffe & Cave, 2002 [1]
        mesophylla Mora-Aguilar & Delgado, 2012 [1]
        metrica Steinheil, 1872 [3]
        miamiensis Howden & Endrödi 1966 [1]
        minuchae Joly, 2003 [1]
        minuta Burmeister, 1847 [1]
        modesta Burmeister, 1847 [4]
        molesta Endrödi, 1969 [1]
        monacha Ratcliffe, 2008 [1]
        monzoni Ratcliffe & Cave, 2009 [1]
        moreti Dechambre, 1992 [2]
        morphoidina Prell, 1937 [1]
        multiplex Casey, 1915 [3]
        munda Kirsch, 1871 [3]
        mustacha Ratcliffe, 2003 [1]
        mutata Harold, 1869 [4]
        nana Dechambre, 1999 [1]
        nigerrima Bates, 1887 [3]
        nigra Dechambre, 1999 [1]
        nigricollis Burmeister, 1847 [3]
        nigritarsis Ratcliffe, 1992 [1]
        nigrobasalis Höhne, 1923 [3]
        nigropicta Dechambre & Endrödi, 1983 [1]
        niguasa Dechambre & Endrodi, 1984 [1]
        nike Ratcliffe, 1992 [1]
        nodanotherwon Ratcliffe, 1992 [1]
        notata (Illiger, 1806) [3]
        obesa Burmeister, 1847 [1]
        obscura Endrödi, 1966 [1]
        obscurata Endrödi, 1966 [1]
        occipitalis Fairmaire, 1892 [1]
        ocellata Burmeister, 1847 [4]
        ochracea Prell, 1937 [1]
        octopunctata Burmeister, 1847 [3]
        ohausiana Höhne, 1923 [4]
        olivieri Arrow, 1911 [1]
        ovulum Bates, 1887 [6]
        pan Ratcliffe, 1992 [1]
        panthera Dechambre, 1979 [1]
        paraflora Martinez, 1978 [3]
        paraguayensis marginella Endrödi, 1966 [1]
        paraguayensis paraguayensis Arrow, 1903 [4]
        parallela (Casey, 1915) [3]
        pardolocarnoi Dechambre, 1995 [1]
        pasadenae mexica Martinez, 1969 [1]
        pasadenae pasadenae (Casey, 1915) [4]
        perconfusa Dechambre, 1992 [1]
        pereirai (Martinez, 1960) [1]
        perforata Arrow, 1913 [4]
        perplexa Ratcliffe, 2008 [1]
        peruana Endrödi, 1966 [1]
        pichinchana Dechambre, 1992 [1]
        picipes (Olivier, 1789) [4]
        picopijola Ratcliffe & Cave, 2006 [1]
        picta Burmeister, 1847 [4]
        pilosa Dupuis, 2006 [1]
        pilosicollis (Saylor, 1936) [1]
        pompanoni Dechambre, 1979 [1]
        poncheli Dechambre, 2005 [3]
        porioni Dechambre, 1979 [4]
        prelli Endrödi, 1967 [1]
        prolongata Arrow, 1902 [3]
        proxima Dechambre, 1997 [1]
        pseudoconfusa Ratcliffe, 1992 [1]
        pseudomelanocephala Dupuis, 1996 [1]
        puberula LeConte, 1863 [1]
        pugnax Arrow, 1914 [4]
        pulchra Dechambre, 1999 [1]
        puncticollis Endrödi, 1966 [1]
        putrida Burmeister, 1847 [4]
        pygidialis Joly, 2000 [1]
        pygidiata Dupuis, 2000 [1]
        quadripunctata Höhne, 1923 [1]
        quatuordecimpunctata Mannerheim, 1829 [1]
        quercina Burmeister, 1847 [4]
        quisqueya Joly, 1998 [1]
        rangelana Chapin, 1935 [1]
        ratcliffei Endrödi, 1977 [1]
        recta Dupuis, 2008 [1]
        regularis (Casey, 1915) [1]
        robusta LeConte, 1863 [1]
        rogezi Dechambre, 1992 [1]
        rondoniana Ratcliffe, 1992 [1]
        rorulenta Höhne, 1923 [1]
        rubescens Bates, 1891 [3]
        rufa Endrödi, 1967 [1]
        rufescens Endrödi, 1967 [1]
        ruficollis Burmeister, 1847 [3]
        rufonigra Demay, 1838 [1]
        rufovaria Arrow, 1911 [4]
        rustica municipalis Höhne, 1923 [1]
        rustica rustica (Olivier, 1789) [4]
        saltini Ratcliffe, 2008 [1]
        sanguinicollis Burmeister, 1847 [3]
        santaritae Ratcliffe, 1992 [1]
        sarahae Ratcliffe, 1992 [1]
        sardadebiae Dechambre, 2005 [3]
        sarpedon Ratcliffe, 1992 [1]
        scarabaeina (Gyllenhal, 1817) [1]
        schmitzorum Ratcliffe, 1992 [1]
        setidiosa LeConte, 1863 [1]
        setosa Burmeister, 1847 [1]
        sexpunctata Laporte de Castelnau 1840 [3]
        signaticollis Burmeister, 1847 [3]
        similis Dechambre, 1980 [3]
        simillima Dechambre, 1999 [1]
        simulatrix Höhne, 1923 [4]
        sinaloae Howden & Endrödi, 1966 [1]
        sinuosa Höhne, 1923 [1]
        sororia Bates, 1887 [3]
        spangleri Joly, 2000 [1]
        sparsa Arrow, 1902 [1]
        spilopyga Erichson, 1847 [3]
        stictica Burmeister, 1847 [4]
        stockwelli Ratcliffe, 2003 [1]
        striata Endrödi, 1963 [1]
        subsignata Burmeister, 1847 [4]
        supernana Dechambre, 1999 [1]
        suturalis Ohaus, 1911 [7]
        sylviae Dechambre, 1995 [1]
        tarsalis Dechambre, 1979 [3]
        testacea Burmeister, 1847 [4]
        tetrica Burmeister, 1847 [3]
        tidula Dechambre, 1999 [1]
        toulgoeti Dechambre, 1992 [4]
        tridentata dominicensis Cartwright & Chalumeau, 1977 [1]
        tridentata tridentata (Fabricius, 1801) [3]
        tronchonii Martinez, 1975 [1]
        tucumana Brèthes, 1904 [1]
        tutilina Burmeister, 1847 [4]
        tylifera Höhne, 1923 [1]
        unamas Ratcliffe, 2003 [1]
        undata (Olivier, 1789) [6]
        unidentata Endrödi, 1980 [4]
        variabilis Burmeister, 1847 [4]
        varians Burmeister, 1847 [3]
        variipenis Dechambre, 1999 [1]
        variolosa Burmeister, 1847 [3]
        verticalis Burmeister, 1847 [3]
        vestita Höhne, 1923 [1]
        vidanoi Dechambre, 1992 [1]
        vidua Endrödi, 1966 [1]
        villosa Blanchard, 1846 [3]
        vincentiae Arrow, 1900 [1]
        vinosa Arrow, 1937 [1]
        virgo Dechambre, 1999 [1]
        viridis Dechambre, 1982 [1]
        virkkii Howden & Endrodi, 1966 [1]
        vittoscutellaris Prell, 1937 [1]
        wandae Hardy, 1974 [1]
        warneri Ratcliffe, 1992 [1]
        weidneri boliviana Dechambre, 1992 [1]
        weidneri weidneri Endrödi, 1964 [5]
        williami Ratcliffe, 1992 [1]
        zischkai Martinez, 1965 [1]
        zodion Ratcliffe, 1992 [1]
        zurstrasseni Endrödi, 1964 [3]
      - DYSCINETUS Harold, 1869 [56]
        australis Joly & Escalona, 2002 [3]
        dubius (Olivier, 1789) [4]
        dytiscoides Arrow, 1911 [3]
        fimosus Herbst, 1789 [1]
        gagates (Burmeister, 1847) [3]
        imitator Ratcliffe, 1986 [1]
        laevicollis Arrow, 1937 [3]
        laevipunctatus Bates, 1887 [4]
        martinezi Joly & Escalona, 2002 [4]
        mendax Joly & Escalona, 2010 [4]
        minor Chapin, 1935 [1]
        morator (Fabricius, 1798) [3]
        olivaceus Höhne, 1923 [3]
        ornaticaudus Ratcliffe, 1986 [1]
        paradytis (Dechambre, 2003) [1]
        picipes picipes (Burmeister, 1847) [3]
        picipes puertoricensis Chalum. 1982 [1]
        plicatus (Burmeister, 1847) [3]
        questeli Chalumeau, 1982 [3]
        rugifrons (Burmeister, 1847) [4]
        sculptus Dupuis, 2006 [1]
        subsericeus (Burmeister, 1847) [1]
      - ERIOSCELIS Burmeister, 1847 [11]
        columbica Endrödi, 1966 [1]
        emarginata (Mannerheim, 1829) [4]
        peruana Saylor, 1946 [1]
        proba (Sharp, 1877) [3]
        sobrina Höhne, 1921 [1]
      - HARPOSCELIS Burmeister, 1847 [3]
        paradoxus Burmeister, 1847 [3]
      - MIMEOMA Casey, 1915 [14]
        acuta (Arrow, 1902) [3]
        englemani Ratcliffe, 1977 [1]
        maculata (Burmeister, 1847) [4]
        nigra Endrödi, 1979 [1]
        signatoides (Höhne, 1923) [4]
      - PELTONOTUS Burmeister, 1847 [33]
        adelphosimilis Jameson & Wada, 2004 [1]
        animus Jameson & Wada, 2009 [1]
        brunnipennis Benderitter, 1934 [1]
        cybele Jameson & Wada, 2009 [1]
        deltamentum Jameson & Wada, 2004 [1]
        favonius Jameson & Wada, 2009 [1]
        fujiokai Jameson & Wada, 2004 [1]
        gracilipodus Jameson & Wada, 2004 [1]
        karubei Muramoto, 2000 [1]
        kyojinus Jameson & Wada, 2004 [1]
        malayensis Arrow, 1910 [2]
        morio Burmeister, 1847 [4]
        mushiyaus Jameson & Wada, 2009 [1]
        nasutus Arrow, 1910 [4]
        nethis Jameson & Wada, 2004 [1]
        podocrassus Jameson & Wada, 2004 [1]
        pruinosus Arrow, 1910 [1]
        rubripennis Miyake & Yamaya, 1994 [1]
        silvanus Jameson & Wada, 2004 [1]
        similis Arrow, 1931 [1]
        sisyrus Jameson & Wada, 2004 [1]
        suehirogarus Jameson & Wada, 2004 [1]
        talangensis Jameson & Jakl, 2010 [1]
        tigerus Jameson & Wada, 2009 [1]
        vittatus Arrow, 1910 [1]
      - RUTELORYCTES Arrow, 1908 [5]
        bis Dechambre, 2006 [1]
        morio (Fabricius, 1798) [3]
      - STENOCRATES Burmeister, 1847 [82]
        agricola Dechambre & Hardy, 2004 [1]
        amazonicus Ratcliffe, 1978 [1]
        ariasi Ratcliffe, 1978 [1]
        batesi Dechambre, 1979 [1]
        beckeri Howden, 1970 [1]
        bicarinatus Robinson, 1947 [4]
        bolivianus Dechambre, 1979 [1]
        bollei Dechambre, 1985 [1]
        canuli Delgado & Castillo, 1991 [1]
        carbo Prell, 1938 [3]
        carinatus Endrödi, 1966 [4]
        celatus Prell, 1938 [3]
        clipeatus Endrödi, 1966 [4]
        cognatus Endrödi, 1966 [1]
        cultor cultor Burmeister, 1847 [3]
        cultor inelegans Arrow, 1913 [1]
        davisorum Endrödi, 1979 [1]
        dubius Endrödi, 1966 [1]
        duplicatus Endrödi, 1967 [1]
        frater Dechambre, 2006 [1]
        haacki Ratcliffe, 1977 [1]
        hardyi Dechambre, 1985 [1]
        hiekei Endrödi, 1967 [1]
        holomelanus (Germar, 1824) [3]
        howdeni Dechambre & Hardy, 2004 [1]
        impeditus Dechambre & Hardy, 2004 [1]
        inpai Ratcliffe, 1978 [1]
        laborator australis Endrödi, 1973 [1]
        laborator laborator (Fabricius, 1775) [1]
        laceyi Ratcliffe, 1978 [1]
        lachaumei Dechambre, 1985 [1]
        laevicollis Kirsch, 1871 [3]
        latus Dechambre, 1979 [1]
        lecourti Dechambre, 2006 [1]
        lichyi Dechambre, 1979 [1]
        ligneus Arrow, 1911 [3]
        mahunkai Endrödi, 1973 [1]
        minutus Endrödi, 1966 [3]
        mollis Endrödi, 1966 [3]
        nasutus Dechambre, 1979 [1]
        omissus Endrödi, 1966 [1]
        pereirai Endrödi, 1969 [1]
        popei Endrödi, 1971 [4]
        porioni Dechambre, 1985 [1]
        pseudoligneus Dechambre & Hardy, 2004 [1]
        rabbanii Ratcliffe, 1977 [1]
        rionegroensis Ratcliffe, 1978 [1]
        rufipennis (Fabricius, 1801) [1]
        rugulosus Endrödi, 1966 [3]
        spinosus Ponchel & Dechambre 2003 [1]
        varzeansis Ratcliffe, 1978 [1]
      - SURUTU Martinez, 1955 [11]
        dytiscoides Martinez, 1955 [3]
        fenni Ratcliffe, 1981 [1]
        hesperius Ratcliffe, 1981 [1]
        jelineki Endrödi, 1975 [1]
        schulzei Endrödi, 1975 [1]
        seabrai Andretta & Martinez, 1956 [3]
    - DYNASTINI [503]
      - ALLOMYRINA ARROW, 1911 [12]
        davidis (Deyrolle, 1878) [4]
        pfeifferi celebensis Silvestre, 1997 [3]
        pfeifferi pfeifferi (Redtenbacher 1867) [4]
      - AUGOSOMA Burmeister, 1841 [8]
        centaurus (Fabricius, 1775) [4]
        hippocrates Miliani, 1995 [3]
      - BECKIUS Dechambre, 1992 [5]
        beccari beccari (Gestro, 1876) [5]
      - CHALCOSOMA Hope, 1837 [44]
        atlas ab. hesperus (Erichson, 1834) [3]
        atlas ab. kirbyi Arrow, 1937 [3]
        atlas atlas (Linnaeus, 1758) [4]
        atlas butonensis Nagai, 2004 [1]
        atlas keyboh Nagai, 2004 [3]
        atlas mantetsu Nagai, 2004 [2]
        atlas simeuluensis Nagai, 2004 [1]
        atlas sintae Nagai, 2004 [3]
        caucassus (Fabricius, 1801) [4]
        caucassus ab. belangeri (Guérin & Méneville, 1834) [3]
        caucassus ab. bidentata Endrödi, 1976 [3]
        caucassus ab. chiron (Olivier, 1789) [3]
        caucassus ab. crassicornis E 1976 [1]
        caucassus ab. edentata Endrödi, 1976 [1]
        caucassus ab. inornata Endrödi, 1976 [1]
        caucassus ab. jansseni Beck, 1937 [2]
        engganensis Nagai, 2004 [1]
        moellenkampi Kolbe, 1900 [4]
      - DYNASTES Kirby, 1825 [57]
        granti Horn, 1870 [5]
        hercules baudrii Pinchon, 1976 [3]
        hercules bleuzeni Silvestre & Dechambre 1995 [4]
        hercules ecuatorianus Ohaus, 1913 [4]
        hercules hercules (Linnaeus, 1758) [4]
        hercules lichyi Lachaume, 1985 [3]
        hercules occidentalis Lachaume, 1985 [3]
        hercules paschoali Grossi & Arnaud, 1993 [3]
        hercules septentrionalis Lachaume, 1985 [4]
        hyllus (Chevrolat, 1843) [4]
        miyashitai Yamaya, 2004 [1]
        maya Hardy, 2003 [3]
        moroni Nagai, 2005 [3]
        neptunus neptunus (Quensel, 1917) [4]
        neptunus rouchei Nagai, 2005 [1]
        satanas Moser, 1909 [3]
        tityus (Linnaeus, 1763) [4]
      - ENDEBIUS Lansberge, 1880 [8]
        florensis florensis (Lansberge, 1879) [1]
        florensis tanimbar Rowland, 2006 [1]
        florensis wetarensis Fujii & Silvestre, 2012 [4]
        mirabilis Silvestre, 2006 [1]
      - EUPATORUS Burmeister, 1847 [31]
        birmanicus Arrow, 1908 [4]
        gracilicornis edai Hirasawa, 1991 [1]
        gracilicornis gracilicornis Arrow, 1908 [4]
        gracilicornis kimioi Hirasawa, 1992 [3]
        hardwickei (Hope, 1831) [4]
        hardwickei ab.cantori (Hope, 1842) [7]
        siamensis (Laporte de Castelnau, 1867) [4]
        sukkiti Miyashita & Arnaud, 1997 [3]
      - GOLOFA Hope, 1837 [95]
        aegeon (Drury, 1770) [4]
        antiquus Arrow, 1911 [1]
        argentinus Arrow, 1911 [4]
        claviger claviger (Linnaeus, 1771) [4]
        claviger puncticollis Thomson, 1860 [3]
        cochlearis Ohaus, 1910 [4]
        costaricensis Bates, 1887 [4]
        eacus Burmeister, 1847 [6]
        gaujoni Lachaume, 1985 [3]
        globulicornis Dechambre, 1975 [4]
        henrypitieri Arnaud & Joly, 2006 [4]
        hirsuta Ratcliffe, 2003 [3]
        imbellis Bates, 1887 [1]
        incas Hope, 1837 [4]
        inermis Thomson, 1859 [1]
        minutus Sternberg, 1910 [3]
        obliquicornis Dechambre, 1975 [3]
        paradoxus Dechambre, 1975 [1]
        pelagon Burmeister, 1847 [4]
        pizarro Hope, 1837 [4]
        porteri Hope, 1837 [4]
        pusilla Arrow, 1911 [4]
        solisi Ratcliffe, 2003 [4]
        spatha Dechambre, 1989 [3]
        tepaneneca Moron, 1995 [1]
        tersander (Burmeister, 1847) [4]
        testudinarius (Prell, 1934) [1]
        unicolor (Bates, 1891) [4]
        wagneri Abadie, 2007 [1]
        xiximeca Moron, 1995 [3]
      - HAPLOSCAPANES Arrow, 1908 [12]
        australicus (Arrow, 1908) [3]
        barbarossa (Fabricius, 1775) [4]
        inermis (Prell, 1911) [1]
        papuanus Dechambre & Drumont, 2004 [3]
      - MEGASOMA Kirby, 1825 [78]
        actaeon actaeon (Linnaeus, 1758) [5]
        actaeon fujitai Nagai, 2003 [1]
        anubis (Chevrolat, 1836) [4]
        cedrosa Hardy, 1972 [3]
        elephas (Fabricius, 1775) [4]
        gyas gyas (Jablonsky, 1785) [4]
        gyas prandii Milani, 2008 [1]
        gyas rumbucheri Fischer, 1968 [5]
        janus Felsche, 1906 [5]
        joergenseni joergenseni (Bruch, 1910) [5]
        joergenseni penyai Nagai, 2003 [1]
        lecontei Hardy, 1972 [1]
        mars (Reiche, 1852) [5]
        nogueirai Moron, 2005 [4]
        occidentalis Bolivar & Jimenez & Martinez, 1963 [5]
        pachecoi Cartwright, 1963 [4]
        punctulatus Cartwright, 1952 [4]
        ramirezorum Silvestre & Arnaud, 2002 [4]
        sleeperi Hardy, 1972 [3]
        svobodaorum Krajcik, 2009 [1]
        thersites LeConte, 1861 [4]
        vogti Cartwright, 1963 [4]
      - PACHYORYCTES Arrow, 1908 [5]
        elongatus Arrow, 1941 [1]
        solidus Arrow, 1908 [3]
      - TRYPOXYLUS Minck, 1920 [26]
        dichotomus dichotomus (Linnaeus, 1771) [6]
        dichotomus politus (Prell, 1934) [4]
        dichotomus septentrionalis (Kôno, 1931) [3]
        dichotomus takarai Kusui, 1976 [3]
        dichotomus tsuchiyai (Nagai, 2006) [1]
        dichotomus tsunobosonis (Kôno, 1931) [4]
        kanamorii (Nagai, 2006) [4]
      - XYLOTRUPES Hope, 1837 [122]
        australicus australicus Thomson, 1859 [4]
        australicus darwinia Rowland, 2006 [1]
        beckeri beckeri Schaufuss, 1885 [4]
        beckeri intermedius Silvestre, 2004 [1]
        carinulus Rowland, 2011 [1]
        clinias buru Rowland, 2011 [1]
        clinias clinias Schaufuss, 1885 [1]
        damarensis Rowland, 2006 [1]
        faber Silvestre, 2002 [3]
        falcatus Minck, 1920 [3]
        gideon borneensis Minck, 1920 [3]
        gideon gideon (Linnaeus, 1767) [3]
        gideon lakorensis Silvestre, 2002 [1]
        gideon sawuensis Silvestre, 2002 [1]
        gideon sondaicus Silvestre, 2002 [4]
        gideon tanahmelayu Rowland, 2006 [1]
        gilleti [3]
        inarmatus Sternberg, 1906 [3]
        lorquini lorquini Schaufuss, 1885 [4]
        lorquini zideki Rowland, 2003 [1]
        lumawigi Silvestre, 2002 [4]
        macleayi asperulus Minck, 1920 [1]
        macleayi macleayi Montrouzier, 1857 [1]
        macleayi szekessyi Endrödi, 1951 [4]
        meridionalis Prell, 1914 [4]
        mirabilis (Silvestre, 2006) [1]
        mniszechi beckeroides Rowland, 2006 [3]
        mniszechi hainaniana Rowland, 2006 [1]
        mniszechi mniszechi Thomson, 1859 [4]
        pachycera Rowland, 2006 [1]
        pauliani dayakorum Silvestre, 2004 [1]
        pauliani pauliani Silvestre, 1997 [3]
        philippinensis peregrinus Rowland, 2006 [1]
        philippinensis philippinensis Endrödi 1957 [3]
        pubescens baumeisteri Schauffus, 1885 [4]
        pubescens beaudeti Silvestre, 2006 [3]
        pubescens gracilis Silvestre, 2006 [1]
        pubescens pubescens Whaterhouse, 1841 [3]
        pubescens sibuyanensis Silvestre, 2006 [1]
        rindaae Fujii, 2011 [1]
        socrates bourgini Paulian, 1945 [3]
        socrates nitidus Silvestre, 2003 [1]
        socrates siamensis Minck, 1920 [3]
        socrates socrates Schaufuss, 1864 [3]
        socrates tonkinensis Minck, 1920 [4]
        striatopunctatus Silvestre, 2003 [3]
        sumatrensis sumatrensis Minck, 1920 [1]
        sumatrensis tanahmelayu Rowland, 2006 [3]
        tadoana Rowland, 2006 [1]
        taprobanus ganesha Silvestre, 2003 [1]
        taprobanus taprobanus Prell, 1914 [3]
        ulysses (Guérin-Méneville, 1830) [5]
        wiltrudae Silvestre, 1997 [1]
    - HEXODONTINI [63]
      - HEMICYRTHUS Reiche, 1860 [13]
        chazeaui Dechambre, 1982 [1]
        costatus Dechambre, 1982 [1]
        elongatus Dechambre, 1982 [1]
        gutierrezi Dechambre, 1982 [1]
        reichei Dechambre, 1982 [1]
        serresii (Montrouzier, 1860) [4]
        villersi (Montrouzier, 1860) [3]
      - HEXODON Olivier, 1789 [44]
        griseosericans Fairmaire, 1901 [3]
        kochi Frey, 1957 [3]
        latissinum Arrow, 1912 [4]
        minutum Sternberg, 1910 [4]
        montandoni Buquet, 1840 [4]
        patella Arrow, 1912 [1]
        quadriplagiatum Frey, 1957 [1]
        reticulatum Olivier, 1789 [5]
        unicolor convexus Decham. 1986 [1]
        unicolor gigas Dechambre, 1986 [1]
        unicolor megaspilosa Decha. 1986 [4]
        unicolor unicolor Olivier, 1789 [4]
        unicostatum unicostatum Arrow, 1912 [4]
        unicostatum simplex Endrödi, 1976 [4]
      - HYBOSCHEMA Péringuey, 1901 [6]
        contractum (Boheman, 1857) [4]
        contractum ab.sobrinum Péringuey 1901 [1]
    - ORYCTINI [851]
      - ANOMACAULUS Fairmaire, 1878 [1]
        fulvovestitus Fairmaire, 1878 [1]
      - BLABEPHORUS Fairmaire, 1898 [3]
        pinguis Fairmaire, 1898 [3]
      - CALYPSORYCTES Howden, 1970 [1]
        dynastoides Howden, 1970 [1]
      - CERATORYCTODERUS Arrow, 1908 [10]
        armatus Dechambre, 2001 [3]
        borchmanni Endrödi, 1959 [1]
        candezei (Snellen van Vollenhoven, 1866) [4]
        mototanii Yamaya, 2011 [1]
      - CLYSTER Arrow, 1908 [28]
        hirokiharukae Yamaya, 2009 [1]
        itys (Olivier, 1789) [4]
        rectus Dechambre, 2002 [1]
        retusus retusus Arrow, 1908 [4]
        retusus nakanei Miyake & Yamaya, 1999 [4]
        rosaemariae Dupuis, 2002 [4]
        scaurus Prell, 1911 [4]
        trachypygus Prell, 1911 [4]
        trinodosus Dechambre, 2002 [1]
      - COELOSIS Hope, 1847 [25]
        bicornis (Leske, 1779) [4]
        biloba (Linnaeus, 1767) [4]
        bourgini (Dechambre, 1976) [1]
        denticornis Arrow, 1937 [4]
        hippocrates Blanchard, 1846 [4]
        inermis Sternberg, 1908 [3]
        sylvanus (Fabricius, 1775) [4]
      - CYPHONISTES Burmeister, 1847 [82]
        arrowi Prell, 1912 [4]
        brachti Prell, 1912 [4]
        burmeisteri Dohrn, 1875 [4]
        camiadei Silvestre, 1999 [1]
        camurus bouyeri Drumont, 1997 [3]
        camurus camurus Karsch, 1881 [4]
        corniculatus corniculatus Burmeister, 1847 [3]
        gasanus Prell, 1912 [4]
        glabricollis (Burmeister, 1847) [4]
        impressicollis Fairmaire, 1898 [3]
        inermicollis Fairmaire, 1894 [5]
        ituriensis Burgeon, 1947 [4]
        latiusculus (Fairmaire, 1887) [4]
        letiranti Hardy, 2003 [1]
        luluensis Burgeon, 1947 [4]
        roeri Endrödi, 1980 [1]
        sublaevis Arrow, 1914 [4]
        tuberculifrons Quedenfeldt, 1884 [4]
        vallatus burgeoni Basilewsky, 1955 [4]
        vallatus deserti Prell, 1912 [4]
        vallatus guineensis Paulian, 1954 [4]
        vallatus nyansae Prell, 1912 [1]
        vallatus septentrionis Prell, 1912 [4]
        vallatus vallatus (Wiedeman, 1823) [3]
      - DICHODONTUS Burmeister, 1847 [29]
        birmanicus Voirin, 1996 [2]
        coronatus Burmeister, 1847 [4]
        crassus Silvestre, 2001 [3]
        croesus (Fabricius, 1801) [4]
        denticornis Voirin, 1996 [1]
        deruptus Prell, 1912 [3]
        fissicornis Arrow, 1911 [1]
        grandis Ritsema, 1882 [4]
        intermedius Dechambre, 1981 [1]
        obesus Prell, 1912 [1]
        pulchellus Prell, 1912 [1]
        punctipennis Arrow, 1937 [3]
      - DINORYCTES Felsche, 1906 [9]
        deguidei Endrödi, 1958 [4]
        truncaticollis (Fairmaire, 1894) [4]
      - ENEMA Hope, 1837 [9]
        endymion Chevrolat, 1843 [4]
        pan (Fabricius, 1775) [4]
      - GIBBORYCTES Endrödi, 1974 [11]
        acuminatus Endrödi, 1978 [1]
        bollei Dechambre, 2006 [1]
        gracilicornis (Prell, 1912) [1]
        szelenyii Endrödi, 1974 [3]
        waldenfelsi (Endrödi, 1977) [4]
      - HETEROGOMPHUS Burmeister, 1847 [112]
        achilles Burmeister, 1847 [4]
        aequatorius (Bates, 1891) [1]
        aidoneus (Perty, 1830) [4]
        amphitryon Burmeister, 1847 [1]
        arrowi Prell, 1912 [1]
        bicuspis Endrödi, 1976 [1]
        binodosus Prell, 1912 [4]
        bispinosus Burmeister, 1847 [3]
        bourcieri Guérin-Méneville, 1851 [4]
        carayoni Dechambre, 1986 [1]
        castaneus Dupuis & Dechambre, 2008 [1]
        chevrolati Burmeister, 1847 [4]
        consanguineus Prell, 1912 [1]
        coriaceus Prell, 1912 [4]
        consors Prell, 1912 [1]
        cribricollis Prell, 1912 [1]
        curvicornis Prell, 1912 [1]
        dejeani Reiche, 1859 [1]
        dilaticollis Burmeister, 1847 [4]
        eteocles Burmeister, 1847 [4]
        flohri (Kolbe, 1906) [3]
        gibbosus Dupuis, Grossi & Grossi, 2012 [1]
        gracilicornis Prell, 1912 [4]
        hiekei Endrödi, 1976 [1]
        hirticollis Prell, 1912 [1]
        hirtus Prell, 1912 [4]
        hopei Burmeister, 1847 [3]
        inarmatus inarmatus Ohaus, 1910 [1]
        inarmatus ochoai Martinez, 1966 [1]
        incornutus Prell, 1912 [1]
        julus Burmeister, 1847 [3]
        laticollis Prell, 1912 [4]
        mirabilis Prell, 1912 [1]
        mniszechi (Thomson, 1859) [4]
        monotuberculatus Dupuis & Dechambre 2008 [3]
        ochoai Martinez, 1966 [1]
        olsoufieffi (Prokofiev, 2012) [1]
        pauson (Perty, 1830) [3]
        peruanus Endrödi, 1976 [1]
        pilosus Dechambre, 1998 [3]
        porioni Dechambre, 1998 [4]
        rubripennis Prell, 1912 [1]
        rugicollis Prell, 1912 [3]
        rugosus (Blanchard, 1846) [1]
        schoenherri Burmeister, 1847 [4]
        telamon (Burmeister, 1847) [1]
        thoas Burmeister, 1847 [3]
        ulysses Burmeister, 1847 [4]
      - HISPANIORYCTES Howden, 1978 [4]
        paulseni Ratcliffe & Cave, 2011 [1]
        rugulopygus Ratcliffe & Cave, 2011 [1]
        sculptilis Ratcliffe & Cave, 2011 [1]
        wittmeri Howden, 1978 [1]
      - HOPLORYCTODERUS Prell, 1933 [14]
        chinensis Voirin, 1996 [1]
        tridens tridens (Montrouzier, 1860) [4]
        tridens caledonicus (Prell, 1914) [4]
        tridens pinorum Paulian, 1991 [4]
      - IRAZUA Ratcliffe, 2003 [1]
        dilicra Ratcliffe, 2003 [1]
      - LICNOSTRATEGUS Prell, 1933 [3]
        endymion (Olivier, 1789) [3]
      - MEGACERAS Hope, 1837 [49]
        bolivianus Dechambre, 1975 [1]
        brevis Dechambre, 1998 [3]
        briansaltini Ratcliffe, 2007 [1]
        celatus Dechambre, 1998 [4]
        crassum Prell, 1914 [4]
        endroedii Dechambre, 1998 [1]
        hoplites Silvestre, 1996 [1]
        inexpectatus Dechambre, 1998 [1]
        inflatum Prell, 1934 [1]
        jason jason (Fabricius, 1775) [4]
        jason stuebeli Kirsch, 1885 [3]
        laevipenne Prell, 1914 [1]
        morpheus Burmeister, 1847 [4]
        pauliani Dechambre, 1975 [1]
        philoctetes (Olivier, 1789) [4]
        porioni Dechambre, 1981 [1]
        quadraticollis Dechambre, 1975 [3]
        remus Reiche, 1859 [3]
        septentrionis angusticollis Ponchel, 1998 [3]
        septentrionis septentrionis Bates, 1887 [4]
      - MEGACEROPSIS Dechambre, 1976 [2]
        lecourti Dechambre, 1996 [1]
        quadridentatus Dechambre, 1976 [1]
      - ORYCTES Illiger, 1798 [224]
        agamemnon agamemnon Burmeister, 1847 [1]
        agamemnon arabicus Fairmaire, 1896 [4]
        agamemnon matthiesseni Reitte, 1907 [4]
        agamemnon sinaicus Walker, 1871 [8]
        amberiensis amberiensis Sternberg, 1910 [1]
        amberiensis viossati Dechambre, 1986 [1]
        amberiensis pauliani Dechambre, 1986 [4]
        anguliceps Fairmaire, 1901 [4]
        ata Semenov & Medvedev, 1932 [3]
        augias (Olivier, 1789) [4]
        boas (Fabricius, 1775) [4]
        borbonicus Dechambre, 1983 [1]
        capucinus Arrow, 1937 [1]
        centaurus Sternberg, 1910 [4]
        cherlonneixi Dechambre, 1996 [4]
        chevrolati Guérin-Méneville, 1844 [1]
        colonicus Coquerel, 1852 [4]
        comoriensis comoriensis Fairmaire, 1893 [3]
        comoriensis intermedius Dechambre, 1986 [6]
        congonis Endrödi, 1969 [4]
        curvicornis Sternberg, 1910 [1]
        elegans Prell, 1914 [4]
        erebus erebus Burmeister, 1847 [6]
        forceps Dechambre, 1980 [4]
        gigas gigas Laporte de Castelnau, 1840 [6]
        gigas insulicola Dechambre, 1986 [3]
        gnu Mohnike, 1874 [4]
        gracilis Prell, 1934 [3]
        heros Endrödi, 1973 [6]
        latecavatus Fairmaire, 1891 [4]
        mayottensis Dechambre, 1983 [4]
        minor Waterhouse, 1876 [1]
        monardi Beck, 1942 [3]
        monoceros (Olivier, 1789) [8]
        nasicornis afghanistanicus Endrödi, 1938 [3]
        nasicornis chersonensis Minck, 1915 [1]
        nasicornis edithae Endrödi, 1938 [1]
        nasicornis grypus Illiger, 1803 [4]
        nasicornis hindenbergi Minck, 1915 [1]
        nasicornis holdhausi Minck, 1914 [1]
        nasicornis illigeri Minck, 1915 [1]
        nasicornis kuntzeni Minck, 1914 [3]
        nasicornis laevigatus Heer, 1841 [4]
        nasicornis latipennis Motschulsky, 1845 [4]
        nasicornis mariei Bourgin, 1949 [4]
        nasicornis nasicornis (Linnaeus, 1712) [4]
        nasicornis ondrejanus Minck, 1916 [3]
        nasicornis polonicus Minck, 1916 [4]
        nasicornis przevalskii Semenov & Medvedev, 1932 [3]
        nasicornis punctipennis Motschulsky, 1860 [4]
        nasicornis shiraticus Endrödi & Petrovitz, 1974 [1]
        nasicornis turkestanicus Minck, 1914 [4]
        nudicauda Arrow, 1910 [1]
        ohausi Minck, 1913 [4]
        owariensis Beauvois, 1805 [4]
        politus Fairmaire, 1901 [1]
        prolixus Wollaston, 1864 [4]
        pyrrhus Burmeister, 1847 [4]
        ranavalo Coquerel, 1852 [4]
        rhinoceros (Linnaeus, 1758) [4]
        richteri Petrovitz, 1958 [1]
        sahariensis de Miré, 1960 [4]
        simiar dujardini Dechambre, 1987 [4]
        simiar simiar Coquerel, 1852 [4]
        sjöstedti Kolbe, 1905 [4]
        tarandus (Olivier, 1789) [4]
        vicinus Gahan, 1900 [3]
      - PODISCHNUS Burmeister, 1847 [13]
        agenor (Olivier, 1789) [4]
        oberthuri Sternberg, 1907 [4]
        sexdentatus (Taschenberg, 1870) [4]
      - SCAPANES Burmeister, 1847 [21]
        affinis Dechambre, 1995 [3]
        australis australis (Boisduval, 1835) [4]
        australis brevicornis Sternberg 1908 [4]
        australis grossepunctatus Sternberg 1908 [4]
        australis salomonensis Sternberg 1908 [5]
      - STRATEGUS Hope, 1837 [98]
        adolescens Kolbe, 1906 [1]
        aenobarbus (Fabricius, 1775) [4]
        ajax (Olivier, 1789) [4]
        aloeus (Linnaeus, 1758) [4]
        anachoreta Burmeister, 1847 [4]
        antaeus (Drury, 1770) [4]
        argentinus Kolbe, 1906 [1]
        atlanticus Ratcliffe, 1976 [1]
        caymani Ratcliffe, 1976 [1]
        centaurus Kolbe, 1906 [4]
        cessus LeConte, 1866 [4]
        craigi Ratcliffe, 1976 [1]
        fallaciosus Kolbe, 1906 [1]
        fascinus Burmeister, 1847 [3]
        hipposiderus Ratcliffe, 1976 [1]
        howdeni Ratcliffe, 1976 [1]
        inermis Arrow, 1947 [1]
        jugurtha Burmeister, 1847 [4]
        longichomperus Ratcliffe, 1976 [4]
        mandibularis Sternberg, 1910 [4]
        moralesdelgadorum Delgado, 1997 [1]
        mormon Burmeister, 1847 [1]
        oblongus (Beauvois, 1805) [6]
        sarpedon (Burmeister, 1847) [4]
        simson (Linnaeus, 1758) [3]
        splendens (Beauvois, 1805) [4]
        surinamensis hirtus Sternberg 1910 [4]
        surinamensis surinamensis Burmeister, 1847 [4]
        symphenax Ratcliffe, 1976 [1]
        syphax (Fabricius, 1775) [4]
        talpa (Fabricius, 1792) [6]
        tarquinius Ratcliffe, 1976 [1]
        temoltzin Moron & Nogueira, 2008 [1]
        validus (Fabricius, 1775) [4]
        verilli Ratcliffe, 1976 [1]
      - TALAUTOCLYSTER Yamaya, 2001 [1]
        marikoae Yamaya, 2001 [1]
      - TEHUACANIA Endrödi, 1975 [1]
        howdeni Endrödi, 1975 [1]
      - TRICHOGOMPHUS Burmeister, 1847 [56]
        acuticollis Arrow, 1908 [1]
        bronchus (Jablonsky, 1785) [4]
        dechambrei Dupuis, 2003 [3]
        excavatus excavatus Mohnike, 1874 [4]
        excavatus fairmairei Arrow, 1911 [3]
        galeatus Dechambre, 1996 [3]
        lunicollis alcides Snellen van Vollenhoven, 1864 [3]
        lunicollis fujiokai Mi. & Yamaya, 1999 [1]
        lunicollis lunicollis Burmeister,1847 [3]
        martabani (Guérin-Méneville, 1834) [4]
        mongol Arrow, 1908 [4]
        pistiloides Voirin, 1996 [1]
        quadridentatus Dechambre, 1981 [1]
        robustus Arrow, 1930 [1]
        rongi Dechambre & Drumont, 2000 [3]
        simson selangorensis Endrödi, 1974 [4]
        simson simson Snellen van Vollenhoven, 1864 [4]
        vicinus Dechambre, 1995 [4]
        zangi Sternberg, 1907 [4]
      - XENODORUS Brême, 1844 [7]
        basquini Silvestre, 1995 [1]
        janus (Fabricius, 1801) [5]
      - XYLORYCTES Hope, 1837 [37]
        corniger Bates, 1887 [3]
        ensifer Bates, 1887 [3]
        furcatus Burmeister, 1847 [4]
        guatemalensis Bitar & Delgado, 2009 [1]
        howdenorum Delgado & Najera, 1992 [4]
        jamaicensis (Drury, 1770) [4]
        lobicollis Bates, 1887 [4]
        splendidus Prell, 1914 [4]
        telephus Burmeister, 1847 [1]
        teuthras Bates, 1887 [4]
        thestalus Bates, 1887 [4]
    - ORYCTODERINI [92]
      - CHALCASTENES Arrow, 1937 [4]
        divinus Endrödi, 1971 [1]
        pulcher Arrow, 1937 [1]
        squamigera (Frey, 1969) [1]
        styracoceros Jameson & Ratcliffe, 2009 [1]
      - CHALCOCRATES Heller, 1903 [15]
        bacchusi Endrödi, 1971 [3]
        borchmanni Endrödi, 1957 [4]
        bretoni Dechambre, 2001 [4]
        felschei Heller, 1903 [2]
        uniformis Heller, 1912 [1]
      - COENORYCTODERUS Prell, 1933 [6]
        candezei (Lansberge, 1880) [4]
        robustus (Prell, 1914) [1]
      - HATAMUS Sharp, 1877 [11]
        gressitti Endrödi, 1971 [4]
        inarmatus Voirin, 1997 [1]
        spinosus Voirin, 1997 [1]
        tarsalis Sharp, 1877 [4]
      - MELANHYPHUS Fairmaire, 1881 [9]
        clypealis (Arrow, 1937) [3]
        nanus (Arrow, 1937) [1]
        philippinus Arrow, 1931 [1]
        platygenioides (Fairmaire, 1881) [3]
      - NEOHYPHUS Heller, 1896 [16]
        kleinschmidti (Fairmaire, 1881) [4]
        philippinus Arrow, 1931 [3]
        semivelutinus celebesus Heller, 1896 [1]
        semivelutinus semivelutinus (Fairmaire, 1883) [4]
        suzukii Yamaya, 2011 [3]
      - ONYCHIONYX Arrow,1914 [7]
        australicus Dechambre, 2003 [1]
        minor Dechambre, 2003 [4]
        opacipennis Arrow, 1914 [1]
      - ORYCTODERINUS Endrödi, 1978 [1]
        walfordorum Endrödi, 1978 [1]
      - ORYCTODERUS Boisduval,1835 [22]
        albertisi Gestro, 1876 [1]
        coronatus Bates, 1877 [3]
        godeffroyi anguliceps Prell, 1934 [4]
        godeffroyi godeffroyi Fairmaire 1877 [4]
        godeffroyi hebridarum Endrödi, 1971 [1]
        gracilior Fairmaire, 1977 [4]
        latitarsis Boisduval 1835 [3]
        sengokui Yamaya, 2010 [1]
      - PARORYCTODERUS Dechambre, 1994 [1]
        cornutus Dechambre, 1994 [1]
    - PENTODONTINI [1741]
      - ACERATUS Prell, 1936 [9]
        davus (Burmeister, 1847) [4]
        phyllis Burmeister, 1847 [4]
      - ADORYPHORUS Blackburn, 1889 [6]
        canei Carne, 1957 [1]
        couloni (Burmeister, 1847) [3]
        mellori Carne, 1957 [1]
      - ALISSONOTUM Arrow, 1908 [49]
        africanum Endrödi, 1974 [1]
        amploides (Endrödi, 1973) [1]
        binodulum (Fairmaire, 1891) [1]
        camerunum (Kolbe, 1905) [3]
        crassum Arrow, 1908 [4]
        cribratellum (Fairmaire, 1893) [4]
        elongatum Arrow, 1910 [1]
        impressicolle Arrow, 1908 [6]
        lerui Dupuis, 1998 [1]
        nitens (Paulian, 1970) [1]
        parvum Zhang, 1984 [1]
        pauper (Burmeister, 1847) [3]
        piceum besucheti Endrödi, 1977 [1]
        piceum fujiokai Miyake & Yamaya 1997 [1]
        piceum krombeini (Endrödi, 1976) [1]
        piceum piceum (Fabricius, 1775) [4]
        pittinoi Endrödi, 1985 [1]
        quadridens Dupuis, 1998 [1]
        rangunense Arrow, 1910 [4]
        simile Arrow, 1910 [4]
        yamayai Drumont, 2013 [4]
      - AMPOTIS Dechambre & Madge, 1979 [4]
        alticola Dechambre & Madge, 1979 [4]
      - ANOMALOMORPHA Arrow, 1908 [6]
        anthracina Arrow, 1908 [3]
        giveni Carne, 1957 [1]
        monteithi Allsopp, 2000 [1]
      - ANORONOTUM Arrow, 1939 [1]
        rufum Arrow, 1939 [1]
      - APHONIDES Rivers, 1889 [6]
        dunnianus (Rivers, 1889) [4]
        turbator Endrödi, 1981 [1]
      - APHONODELUS Kolbe, 1905 [6]
        coriaceipennis Kolbe, 1905 [1]
        mraci Dechambre, 1993 [4]
      - APHONUS LeConte, 1856 [19]
        brevicruris Cartwright, 1944 [2]
        castaneus (Melsheimer, 1845) [3]
        densicauda Casey, 1915 [3]
        texanus Gill & Howden, 1985 [4]
        tridentatus (Say, 1823) [3]
        variolosus (LeConte, 1847) [3]
      - AUSTRALOCUS Dupuis, 2010 [1]
        bicornutus Dupuis, 2010 [1]
      - BARUTUS Ratcliffe, 1981 [3]
        hartmanni Ratcliffe, 1981 [3]
      - BOTHYNUS Hope, 1837 [95]
        alvarengai Martinez, 1983 [3]
        ascanius (Kirby, 1818) [4]
        complanus (Burmeister, 1847) [4]
        complexus Dechambre, 2006 [3]
        cribrarius (Fairmaire, 1878) [2]
        cunctator (Mannerheim, 1829) [4]
        cyclops (Burmeister, 1847) [3]
        cylindricus (Arrow, 1937) [4]
        dasypleurus (Germar, 1824) [6]
        deiphobus (Burmeister, 1847) [4]
        herteli Endrödi, 1968 [4]
        entellus (Serville, 1828) [4]
        exaratus (Burmeister, 1847) [5]
        herbivorus (Arrow, 1937) [4]
        horridus Endrödi, 1968 [4]
        laevipennis Arrow, 1937 [4]
        lancifer Dechambre, 1981 [1]
        laticifex Burmeister, 1847 [3]
        medon (Germar, 1824) [4]
        minor Steinheil, 1872 [3]
        niger Blanchard, 1846 [1]
        nyx Ratcliffe, 2010 [3]
        peninsularis Casey, 1915 [1]
        quadridens (Taschenberg, 1870) [3]
        ruginasus LeConte, 1856 [1]
        stenelus (Burmeister, 1847) [3]
        striatellus (Fairmaire, 1878) [3]
        tricornis Arrow, 1937 [1]
        validus (Burmeister, 1847) [1]
        villiersi Endrödi, 1968 [4]
      - CALICNEMIS Laporte de Castelnau, 1832 [13]
        atlanticus Mosconi, 1996 [1]
        bahilloi Lopez-colon, 2003 [1]
        bercedoi Lopez-Colon, 2004 [1]
        latreillei Laporte de Castelnau, 1832 [1]
        obesa Erichson, 1841 [4]
        sardiniensis Leo, 1985 [4]
      - CALLISTEMONUS Péringuey, 1901 [1]
        intrusus Péringuey, 1901 [1]
      - CARNEIOLA Endrödi, 1974 [1]
        zealandica Endrödi, 1974 [1]
      - CARNEODON Ozdikmen, 2009 [16]
        bidens (Blackburn, 1896) [1]
        glauerti (Carne, 1957) [1]
        intermedius (Blackburn, 1896) [1]
        laevicollis (MacLeay, 1871) [1]
        laevipennis (Blackburn, 1896) [3]
        laevis (Burmeister, 1847) [1]
        meyricki (Blackburn, 1896) [1]
        occidentalis (MacLeay, 1888) [1]
        pecuarius (Reiche, 1860) [4]
        simplex (Carne, 1957) [1]
      - CARNEORYCTES Ozdikmen, 2009 [21]
        ater (Lea, 1917) [1]
        brittoni (Carne, 1957) [1]
        griseopilosus (Lea, 1917) [1]
        minchami (Carne, 1981) [1]
        monstrosus (Blackburn, 1895) [1]
        nigripennis (Lea, 1917) [1]
        peterseni (Endrödi, 1967) [1]
        pimbus (Carne, 1957) [1]
        psilus (Carne, 1957) [1]
        semicalvus (Lea, 1917) [1]
        sulcatus (Arrow, 1914) [1]
        tectus (Blackburn, 1891) [3]
        tricornutus (Howden & Maly, 2005) [1]
        trifidus (Blackburn, 1895) [3]
        truncatus (Carne, 1957) [1]
        wingarus (Carne, 1957) [1]
      - CAVONUS Sharp, 1875 [16]
        acutifrons Lea, 1917 [1]
        armatus Sharp, 1875 [9]
        bidens (Blackburn, 1896) [1]
        leai Carne, 1957 [1]
        niger (Blackburn, 1888) [1]
        sculpturatus Blackburn, 1888 [1]
        sharpi Blackburn, 1888 [1]
      - CHEIROPLATYS Hope, 1837 [37]
        amphioxus Arrow, 1937 [1]
        bifossus Blackburn, 1896 [4]
        borealis Carne, 1957 [3]
        carnei Endrödi, 1971 [1]
        castaneus Lea, 1917 [1]
        excavatus Lea, 1917 [4]
        inconspicuus MacLeay, 1887 [3]
        laevicollis Arrow, 1937 [4]
        latipes (Guérin-Méneville, 1830) [6]
        nitidus Carne, 1957 [4]
        solidus Carne, 1957 [1]
        volsellus Carne, 1976 [4]
      - COLLAGENUS Hardy & Ratcliffe, 2004 [1]
        dasysternus Hardy & Ratcliffe, 2005 [1]
      - COPTOGNATHUS Burmeister, 1847 [9]
        attila (Ménétriès, 1849) [4]
        crassipes Burmeister, 1847 [4]
      - CORYNOPHYLLUS Hope, 1845 [25]
        andersoni Blackburn, 1888 [1]
        antennalis Dechambre, 2003 [1]
        arrowi Carne, 1957 [1]
        curvicornis Lea, 1919 [1]
        debilis Fairmaire, 1877 [1]
        deserti Carne, 1957 [1]
        dorrigoensis Carne, 1957 [1]
        elricus Carne, 1957 [1]
        fortnumi Hope, 1845 [5]
        interocularis Lea, 1919 [1]
        macdougalli Carne, 1957 [1]
        major Sharp, 1873 [3]
        modestus Blackburn, 1888 [3]
        pygidialis Carne, 1957 [1]
        teinogenoides Carne, 1957 [1]
        tulanus Carne, 1957 [1]
      - COSCINOCEPHALUS Prell, 1936 [6]
        cribrifrons (Schaeffer, 1906) [4]
        tepehuanus Moron & Ratcliffe, 1996 [1]
      - DALGOPUS Dechambre, 2001 [8]
        aethiopicus Dechambre, 2001 [4]
        major Dechambre, 2001 [3]
      - DASYGNATHUS MacLeay, 1819 [16]
        blattocomes Carne, 1978 [4]
        dejeani MacLeay, 1819 [4]
        globosus Arrow, 1914 [4]
        trituberculatus Blackburn, 1889 [3]
      - DENHEZIA Dechambre, 2006 [1]
        caucacola Dechambre, 2006 [1]
      - DILOBODERUS Reiche, 1859 [4]
        abderus (Sturm, 1826) [4]
      - DIPELICUS Hope, 1845 [104]
        acutus Silvestre, 2006 [3]
        alveolatus (Heller, 1897) [4]
        bicarinatus Silvestre, 2006 [1]
        bidens Arrow, 1910 [1]
        borneensis Arrow, 1911 [1]
        bovilli (Blackburn, 1888) [7]
        cantator Arrow, 1910 [1]
        cantori (Hope, 1842) [4]
        centratus Endrödi, 1969 [4]
        daedalus (Fabricius, 1792) [3]
        deiphobus deiphobus (Sharp, 1873) [4]
        deiphobus loi Kobayashi, 1995 [3]
        deiphobus robustus (Heller, 1897) [3]
        duplex (Sharp, 1875) [4]
        fastigatoides Prokofiev, 2012 [4]
        fastigatus Endrödi, 1969 [3]
        furcatus Endrödi, 1971 [1]
        geryon (Drury, 1770) [1]
        hircus (Fabricius, 1775) [3]
        indicus Endrödi, 1969 [1]
        integriceps integriceps (Fairmaire, 1877) [4]
        integriceps monotuberculatus Miyake & Yamaya, 1997 [1]
        integriceps ritsemae (Heller, 1897) [1]
        lacordairei (Sharp, 1873) [4]
        minusculus Voirin, 1996 [1]
        mitratus Silvestre, 2010 [4]
        montrouzieri montrouzieri (Montrouzier, 1860) [4]
        montrouzieri oryctoides (Fairmaire, 1883) [1]
        montrouzieri salomonensis Arrow [1]
        nasutus Bates, 1877 [1]
        optatus (Sharp, 1875) [4]
        pyramidalis Silvestre, 2006 [1]
        quadratifer (Heller, 1897) [3]
        schultzjenai Silvestre, 2007 [1]
        sinuosus Silvestre, 2006 [4]
        triangularis nasutus Bates, 1877 [1]
        triangularis triangularis (Montrouzier, 1855) [7]
        tuberosus Silvestre, 2012 [4]
      - EBOLOWANIUS Dechambre, 1979 [1]
        mirei Dechambre, 1979 [1]
      - ENAROTADIUS Endrödi, 1971 [4]
        gressitti Endrödi, 1971 [4]
      - ENDROEDIANIBE Chalumeau, 1981 [2]
        bozzolai Chalumeau, 1981 [1]
        xerozoyphius Ratcliffe & Cave, 2011 [1]
      - ENRACIUS Dechambre, 1999 [1]
        longipes Dechambre, 1999 [1]
      - EPIRONASTES Carne, 1957 [3]
        abruptus Carne, 1957 [1]
        limbatus Carne, 1957 [1]
        nigrisetosus Carne, 1957 [1]
      - EREMOBOTHYNUS Ohaus, 1910 [3]
        bicuspis Endrödi, 1969 [1]
        cornutus (Steinheil, 1872) [1]
        hirsutus Grossi & Abadie 2008 [1]
      - EUCOPIDOCAULUS Prell, 1912 [7]
        biintrusa (Heller, 1913) [1]
        platypygus Prell, 1912 [1]
        tridentipes (Arrow, 1911) [4]
      - EUETHEOLA Bates, 1888 [18]
        bidentata (Burmeister, 1847) [4]
        hippocrepis Prokofiev, 2012 [1]
        humilis (Burmeister, 1847) [4]
        latipennis Arrow, 1911 [3]
        sibericana Prokofiev, 2012 [1]
        subglabra (Schaeffer, 1909) [4]
      - EUTYCTUS Semenov, 1889 [3]
        deserti Semenov, 1889 [3]
      - GILLASPYTES Howden, 1980 [1]
        janzeni Howden, 1980 [1]
      - GORDITUS Ratcliffe, 2010 [1]
        rotundus Ratcliffe, 2010 [1]
      - HAPLOSOMA Semenov, 1889 [3]
        ordinatum Semenov, 1889 [3]
      - HETEROCONUS Kolbe, 1900 [15]
        excellens (Arrow, 1941) [1]
        fairmairei Dechambre, 1976 [1]
        major Dechambre, 1976 [1]
        muticus (Burmeister, 1847) [3]
        nanus (Blancsik, 1892) [1]
        paradoxus Endrödi, 1968 [3]
        pauliani Dechambre, 1986 [1]
        peyrierasi Dechambre, 1976 [3]
      - HETEROGLOBUS Dupuis & Dechambre, 2008 [1]
        obesus Dupuis & Dechambre, 2008 [1]
      - HETEROLIGUS Kolbe, 1900 [44]
        appius (Burmeister, 1847) [6]
        bianoensis Burgeon, 1947 [4]
        brevis (Quedenfeldt, 1888) [1]
        fulvopilosus Prell, 1914 [1]
        geotrupinus (Quedenfeldt, 1884) [3]
        meles (Billberg, 1815) [4]
        minor Burgeon, 1947 [3]
        miser Arrow, 1941 [1]
        pentodontinus (Kolbe, 1905) [3]
        peringueyi Arrow, 1911 [3]
        planifrons Prell, 1914 [3]
        rojkoffi Dupuis, 2010 [3]
        sebastiani Endrödi, 1973 [1]
        truncaticeps (Fairmaire 1887) [4]
        vagabundus (Arrow, 1914) [3]
      - HETERONYCHUS Dejean, 1833 [201]
        abyssinicus Jack, 1923 [3]
        amplipennis Benderitter, 1915 [3]
        amploides Endrödi, 1973 [1]
        amplus Kolbe, 1900 [3]
        andersoni Jack, 1923 [1]
        annulatus Bates, 1891 [3]
        approximans Kolbe, 1900 [3]
        arator arator (Fabricius, 1775) [4]
        arator australis Endrödi, 1961 [3]
        arator borealis Endrödi, 1961 [4]
        arator centralis Endrödi, 1961 [4]
        arator clypealis Fairmaire, 1893 [1]
        arator occidentalis Endrödi, 1961 [1]
        arator rugifrons Fairmaire, 1871 [4]
        ascanius Kolbe, 1900 [1]
        atratus Klug, 1855 [3]
        basilewskyi Endrödi, 1962 [4]
        bituberculatus Kolbe, 1900 [3]
        brittoni Ferreira, 1965 [4]
        carvalhoi Ferreira, 1970 [1]
        citernii Paoli, 1934 [4]
        congoensis Kolbe, 1900 [4]
        consimillis Kolbe, 1900 [3]
        cordatus Endrödi, 1974 [3]
        costatus Lansberge, 1886 [1]
        cricetus (Hausmann, 1807) [3]
        curtulus Fairmaire, 1887 [3]
        desaegeri Burgeon, 1947 [4]
        digitiformis Zhang, 1983 [3]
        fossor Reiche, 1847 [3]
        impudens Jack, 1923 [4]
        infans Kolbe, 1897 [1]
        inoportunus Ferreira, 1966 [1]
        insignificus gracilis Paulian, 1954 [1]
        insignificus insignificus Jack, 1923 [3]
        intermedius Zhang, 1983 [3]
        jacki Arrow, 1936 [1]
        krombeini Endrödi, 1976 [1]
        licas (Klug, 1835) [4]
        lioderes Redtenbacher, 1867 [4]
        lusingae Endrödi, 1974 [4]
        minimus Dupuis, 2012 [1]
        minutus Burmeister, 1847 [3]
        mollis Endrödi, 1961 [4]
        monodi Paulian, 1954 [3]
        mosambicus Péringuey, 1908 [4]
        muticus Benderitter, 1915 [3]
        paolii Arrow, 1936 [4]
        parumpunctatus Burmeister, 1847 [1]
        parvus Burmeister, 1847 [7]
        pauperatus Péringuey, 1901 [4]
        plebejus (Klug, 1832) [3]
        puerilis Kolbe, 1900 [3]
        puncticollis Jack, 1923 [4]
        punctolineatus borealis Endrödi, 1969 [1]
        punctolineatus punctolineatus Fairmaire, 1893 [3]
        pygidialis Kolbe, 1900 [4]
        rusticus niger (Klug, 1855) [4]
        rusticus rusticus (Klug, 1832) [3]
        sabackyi Endrödi, 1975 [1]
        sacchari Arrow, 1908 [4]
        similis Dechambre, 1992 [4]
        simulans Jack, 1923 [3]
        sublaevis (Fairmaire, 1891) [3]
        tenuestriatus Fairmaire, 1893 [3]
        tesari Endrödi, 1968 [1]
        tristis Boheman, 1857 [3]
        vixstriatus Jack, 1923 [3]
        wittei Janssens, 1942 [4]
      - HETEROSTRIATUS Dupuis, 2006 [1]
        lerui Dupuis, 2006 [1]
      - HIEKEIANUS Endrödi, 1978 [1]
        tuberculatus Endrödi, 1978 [1]
      - HOMOEOMORPHUS Burmeister, 1847 [2]
        nitidus Dechambre & Ponchel, 1999 [1]
        rufipes Burmeister, 1847 [1]
      - HYLOBOTHYNUS Ohaus, 1910 [9]
        arnaudi Dechambre, 1979 [4]
        columbianus Endrödi, 1969 [1]
        marielae Abadie, 2011 [1]
        obesus Ohaus, 1910 [1]
        pontis Ratcliffe, 1981 [1]
      - HYPHORYCTES Blackburn, 1895 [4]
        maculatus Blackburn, 1895 [4]
      - IDIOSCHEMA Arrow, 1914 [1]
        karruense Arrow, 1914 [1]
      - INDIERALIGUS Deschambre, 1979 [2]
        bozzolai (Chalumeau, 1981) [1]
        michelii Dechambre, 1979 [1]
      - LIGYRUS Burmeister, 1847 [85]
        adoceteus Ratcliffe & Cave, 2010 [1]
        bidentulus Fairmaire, 1892 [1]
        bituberculatus (Palisot de Beauvois, 1805) [4]
        burmeisteri Steinheil, 1872 [4]
        cicatricosus Prell, 1937 [1]
        cuniculus (Fabricius, 1801) [4]
        discrepans Escalona & Joly, 2006 [1]
        ebenus (De Geer, 1774) [4]
        fossor (Latreille, 1833) [3]
        gibbosus gibbosus (De Geer, 1774) [4]
        gibbosus morio (LeConte, 1847) [1]
        glanucai Dechambre & Lumaret, 1985 [1]
        gyas (Erichson, 1848) [4]
        laevicollis Bates, 1887 [4]
        maimon (Erichson, 1847) [4]
        maternus Prell, 1937 [3]
        nasutus (Burmeister, 1847) [4]
        neglectus (LeConte, 1847) [3]
        peruanus Endrödi, 1970 [1]
        pullus Prell, 1937 [1]
        pumilus Prell, 1937 [1]
        relictus (Say, 1825) [4]
        rosettae Endrödi, 1968 [4]
        rubripes (Boheman, 1858) [3]
        sallaei Bates, 1887 [4]
        selanderi (Cartwright, 1959) [4]
        similis Endrödi, 1968 [4]
        subtropicus Blatchley, 1922 [3]
        villosus (Burmeister, 1847) [4]
      - LONCHOTUS Burmeister, 1847 [44]
        aldabrensis Dechambre, 1986 [1]
        ambrensis Dechambre, 1986 [4]
        astovensis Arrow, 1911 [1]
        crassus (Klug, 1832) [8]
        endroedii Dechambre, 1986 [3]
        latus Arrow, 1911 [4]
        lentus Burmeister, 1847 [4]
        minor Dechambre, 1986 [1]
        mohelicus Paulian, 1960 [4]
        politus Arrow, 1908 [4]
        punctatissimus Arrow, 1908 [4]
        rotundus Arrow, 1911 [1]
        rugosicollis Sternberg, 1910 [3]
        sicardi Arrow, 1911 [1]
      - MARRONUS Coquerel, 1866 [3]
        borbonicus Coquerel, 1866 [3]
      - MELLISSIUS Wollaston, 1869 [7]
        adumbratus Wollaston, 1875 [1]
        eudoxus Wollaston, 1877 [3]
        oryctoides Decelle, 1972 [1]
        popei Endrödi, 1972 [1]
      - METANASTE Arrow, 1911 [15]
        australis (Fauvel, 1862) [3]
        bicornis bicornis Lea, 1919 [1]
        bicornis papua Endrödi, 1971 [1]
        irregularis (MacLeay, 1871) [1]
        picipes (MacLeay, 1871) [4]
        vulgivagus (Olliff, 1889) [4]
      - MICRORYCTES Arrow, 1908 [15]
        apicalis Arrow, 1908 [1]
        confinis Zhang, 1991 [4]
        hainanensis Zhang, 2002 [1]
        kanarensis Arrow, 1908 [1]
        katagirii Yamaya, 2000 [1]
        monodon (Fairmaire, 1893) [4]
        silvestrei Drumont, Dechambre, Yamaya & Verdugo, 2011 [2]
      - MUSURGUS Vauloger, 1898 [5]
        sabulicola Dechambre & Pope, 1980 [1]
        stridens Vauloger, 1898 [3]
      - NEATOCNEMIS Prell, 1936 [2]
        minor Carne, 1957 [1]
        punctata Sharp, 1875 [1]
      - NEOCNECUS Sharp, 1878 [2]
        minor (Carne, 1957) [1]
        punctata (Sharp, 1875) [1]
      - NEOCORYNOPHYLLUS Carne, 1985 [1]
        aputaringus Carne, 1985 [1]
      - NEODASYGNATHUS Carne, 1957 [7]
        corynophylloides Carne, 1957 [1]
        davidsoni Carne, 1957 [1]
        juba (Kirby, 1818) [4]
      - NEOMETANASTES Yamaya, 2010 [1]
        hidakai Yamaya, 2010 [1]
      - NEONASTES Carne, 1957 [3]
        glabricollis (MacLeay, 1887) [1]
        uptoni Carne, 1985 [1]
      - NEORYCTES Arrow, 1908 [9]
        galapagoensis (Waterhouse, 1845) [1]
        linelli Mutchler, 1938 [1]
        moreti Dechambre, 1991 [3]
        williamsi Cook, Howden & Peck, 1995 [3]
      - NEPHRODOPUS Sharp, 1873 [4]
        enigma Sharp, 1873 [4]
      - NIMBACOLA Paulian, 1970 [4]
        lamottei Paulian, 1970 [4]
      - NOVAPUS Sharp, 1875 [19]
        adelaidae Blackburn, 1888 [4]
        bifidus Lea, 1910 [1]
        carnei Endrödi, 1974 [1]
        crassus Sharp, 1875 [3]
        laticollis Blackburn, 1890 [1]
        macfarlandi Carne & Allsopp, 1987 [1]
        nitidus Blackburn, 1895 [1]
        obscurus (MacLeay, 1871) [1]
        parvus Lea, 1919 [1]
        rugosicollis Blackburn, 1890 [2]
        simplex Sharp, 1875 [1]
        undarus Carne, 1957 [1]
      - ORIZABUS Fairmaire, 1878 [48]
        amalgamatus Ratcliffe & Cave, 2010 [1]
        australis Ratcliffe & Solis, 2013 [1]
        batesi Prell, 1914 [4]
        botox Ratcliffe & Cave, 2006 [1]
        brevicollis Prell, 1914 [1]
        clunalis (LeConte, 1856) [4]
        cuernavacensis Delgado & D. 1990 [3]
        dechambrei Moron, Tapia. 2003 [1]
        delgadoi Ratcliffe & Cave, 2010 [1]
        endrödianus Moron, 1981 [1]
        epithecus Ratcliffe & Cave, 2010 [1]
        fairmairei (Bates, 1887) [1]
        hondurensis Ratcliffe & Cave, 2006 [1]
        isodonoides Fairmaire, 1878 [4]
        ligyroides Horn, 1885 [4]
        mcclevei Warner, 2011 [1]
        mezclus Ratcliffe & Cave, 2010 [1]
        puchicus Ratcliffe & Cave, 2006 [1]
        pyriformis (LeConte, 1847) [4]
        ratcliffei Delgado 2008 [3]
        rawlinsi Dechambre, 1993 [1]
        rubricollis Prell, 1914 [1]
        subaziro Ratcliffe, 1994 [2]
        teamscaraborum Ratcliffe & Cave, 2006 [1]
        thomasi Ratcliffe & Cave, 2010 [1]
        tuberculatus Prell, 1914 [1]
        vulcanicus Moron, Tapia. 2003 [1]
      - ORSILOCHUS Burmeister, 1847 [7]
        cornutus (Thunberg, 1818) [3]
        kochi Ferreira, 1965 [3]
      - ORTHOCAVONUS Carne, 1957 [1]
        occidentalis (Blackburn, 1888) [1]
      - ORYCTOMORPHUS Guérin-Méneville, 1830 [6]
        bimaculatus Guérin-Méneville, 1830 [1]
        laevipennis Germain, 1855 [3]
        maculicollis Guérin-Méneville, 1838 [1]
      - OXYGRYLIUS Casey, 1915 [9]
        peninsularis Casey, 1915 [4]
        ruginasus (LeConte, 1856) [4]
      - OXYLIGYRUS Arrow, 1908 [18]
        acutus Endrödi, 1966 [1]
        cayennensis Endrödi, 1972 [3]
        contractus Dupuis, 2010 [1]
        larssoni Endrödi, 1969 [1]
        nasutus (Burmeister, 1847) [1]
        peruanus Endrödi, 1966 [3]
        politus Ohaus, 1910 [1]
        rostratus (Burmeister, 1847) [1]
        spitzi Endrödi, 1972 [1]
        zoilus (Olivier, 1789) [4]
      - PAPUANA Arrow, 1911 [85]
        angustus Arrow, 1914 [4]
        armicollis (Fairmaire, 1883) [3]
        biroi Endrödi, 1969 [4]
        cheesmanae Arrow, 1941 [1]
        cognata Endrödi, 1969 [1]
        coronata Endrödi, 1971 [1]
        fascicula Voirin, 1997 [1]
        fortepunctata Heller, 1913 [3]
        hardyi Endrödi, 1971 [1]
        huebneri fallax Prell, 1934 [1]
        huebneri huebneri (Fairmaire, 1879) [4]
        inermis Prell, 1912 [1]
        japenensis Arrow, 1941 [4]
        lansbergei badia (Prell, 1912) [4]
        lansbergei lansbergei (Schaufuss, 1887) [4]
        matsumotoi Yamaya, 2009 [1]
        miyakei Yamaya, 2006 [6]
        oryctoides Dechambre, 2005 [4]
        philippinica Arrow, 1937 [3]
        sedlaceki Endrödi, 1971 [1]
        splendens Prell, 1912 [4]
        sumbana Miyake & Yamaha, 1999 [1]
        sumbawana Miyake & Yamaya, 1999 [1]
        szentivanyi Endrödi, 1971 [1]
        tibialis Arrow, 1941 [4]
        timorensis Miyake & Yamaya, 1999 [1]
        trinodosa Prell, 1912 [4]
        uniformis (Prell, 1934) [3]
        uninodis Prell, 1912 [4]
        woodlarkiana laevipennis Arrow, 1911 [4]
        woodlarkiana tenuistriata (Aulmann, 1911) [1]
        woodlarkiana woodlarkiana (Montrouzier, 1855) [4]
      - PARANODON Cockerell, 1905 [4]
        coquereli (Fairmaire, 1871) [4]
      - PARAPUCAYA Prell, 1934 [6]
        amazonica Prell, 1934 [1]
        nodicollis (Kirsch, 1873) [4]
      - PARETERONYCHUS Prell, 1934 [1]
        dyscinetoides Prell, 1934 [1]
      - PARISOMORPHUS Schaufuss, 1890 [14]
        bouvieri (Fairmaire, 1899) [1]
        descarpentriesi (Lebis, 1961) [3]
        ganglaueri (Fairmaire, 1901) [1]
        hilaris Schaufuss, 1890 [4]
        niger Dechambre, 1976 [3]
        viettei Dechambre, 1986 [1]
      - PENTODINA Endrödi, 1968 [1]
        peruviana Endrödi, 1968 [1]
      - PENTODON Hope, 1837 [75]
        algerinus algerinus (Herbst, 1789) [4]
        algerinus bispinifrons Reitter, 1894 [3]
        algerinus dispar Baudi, 1870 [3]
        algerinus indicus Endrödi, 1967 [1]
        algerinus tchadense Paulian, 1954 [4]
        bengalense Arrow, 1910 [4]
        bidens bidens (Pallas, 1771) [4]
        bidens punctatus (Villers, 1789) [4]
        bidens sulcifrons Küster, 1848 [3]
        caminarius caminarius (Faldermann, 1835) [4]
        caminarius elatus Kuster, 1847 [1]
        distandidens distandidens Reitter, 1898 [3]
        distandidens gobicum Endrödi, 1965 [1]
        idiota gumariensis Petrovitz, 1961 [1]
        idiota idiota (Herbst, 1789) [4]
        idiota mennon Jakovlev, 1902 [3]
        insularis Zhang, 1991 [1]
        intermedius Voirin, 1997 [1]
        kuwaitense Endrödi & Al-Houty, 1985 [1]
        millini Miksic, 1962 [1]
        minutus Reitter, 1887 [3]
        nasutus Petrovitz, 1964 [1]
        quadridens bidentulus (Fairmaire, 1887) [3]
        quadridens mongolicum Motschulsky, 1849 [4]
        quadridens patruelis Frivaldszky, 1890 [1]
        quadridens quadridens Gebler, 1845 [1]
        topali Endrödi, 1974 [3]
        variolopunctatus deserti (Heyden, 1899) [3]
        variolopunctatus variolopunctatus Fairmaire, 1879 [4]
      - PENTODONTOSCHEMA Péringuey, 1901 [12]
        caper Prell, 1936 [4]
        capicola Péringuey, 1901 [1]
        deceptor Péringuey, 1901 [1]
        diversum Péringuey, 1901 [3]
        endrodyyoungai Endrödi, 1975 [1]
        nireum (Burmeister, 1847) [1]
      - PERICOPTUS Burmeister, 1847 [13]
        frontalis Broun, 1904 [1]
        nitidulus Broun, 1880 [1]
        punctatus (White, 1846) [3]
        studipus Sharp, 1878 [3]
        truncatus (Fabricius, 1787) [4]
      - PHILOSCAPTUS Brèthes, 1919 [6]
        bonariensis (Burmeister, 1847) [4]
        laminiger Dechambre, 1979 [1]
      - PHYLLIOCEPHALA Blackburn, 1890 [2]
        interioris Blackburn, 1894 [1]
        nigrohirta Blackburn, 1890 [1]
      - PHYLLOGNATUS Eschscholtz, 1830 [18]
        burmeisteri Arrow, 1911 [4]
        dionysius (Fabricius, 1792) [4]
        excavatus Forster, 1771 [4]
        orion (Olivier, 1789) [4]
        sabatinelli Endrödi, 1979 [1]
      - PIMELOPUS Erichson, 1842 [23]
        dubius dubius Blackburn, 1888 [4]
        dubius occidentalis Carne, 1957 [1]
        fisheri (Montrouzier, 1860) [4]
        minor Carne, 1957 [1]
        noctis (Olliff, 1889) [1]
        nothus Burmeister, 1847 [4]
        porcellus crassus Blackburn, 1888 [1]
        porcellus decipiens Blackburn, 1905 [1]
        porcellus porcellus Erichson, 1847 [4]
        similis Voirin, 1997 [1]
      - PODALGUS Burmeister, 1847 [10]
        cuniculus arabicus (Fairmaire, 1895) [1]
        cuniculus cuniculus Burmeister, 1847 [3]
        cuniculus indicus Endrödi, 1966 [1]
        cuniculus infantulus (Semenov, 1890) [3]
        cuniculus reichei Paulian, 1954 [1]
      - PRIONORYCTES Arrow, 1911 [26]
        breviusculus Arrow, 1911 [3]
        camerunus Prell, 1914 [4]
        capreolus (Quedenfeldt, 1884) [6]
        intermedius Prell, 1912 [4]
        laticollis Prell, 1912 [4]
        rufopiceus Arrow, 1911 [3]
        validus Lea, 1917 [1]
      - PSEUDOCAVONUS Blackburn, 1890 [1]
        antennalis Blackburn, 1890 [1]
      - PSEUDOHOMONYX Arrow, 1908 [32]
        apoensis Miyake & Yamaya, 1997 [1]
        belitungensis Miyake & Yamaya, 1997 [1]
        borneensis Arrow, 1908 [4]
        crassus Arrow, 1911 [4]
        extradentatus Miyake & Yamaya, 1997 [1]
        fuziokai Miyake & Yamaya, 1998 [1]
        javanus javanus (Burmeister, 1847) [3]
        javanus sumatrensis (Fairmaire, 1896) [4]
        luzonicus Miyake & Yamaya, 2000 [1]
        mindroensis Miyake & Yamaya, 2000 [1]
        negros Miyake & Yamaya, 1997 [4]
        palavanus Prell, 1912 [1]
        samarensis Miyake & Yamaya, 1997 [4]
        similis Miyake & Yamaya, 1999 [1]
      - PSEUDORYCTES Sharp, 1873 [12]
        bidentifrons Lea, 1926 [1]
        dispar Sharp, 1875 [1]
        etunus Carne, 1957 [1]
        mullerianus (White, 1859) [1]
        semicrudus Lea, 1917 [1]
        storeyi Carne, 1980 [1]
        turritus (MacLeay, 1888) [1]
        validus Lea, 1917 [3]
        wilsoni Carne, 1957 [1]
      - PUCAYA Ohaus, 1910 [9]
        castanea Ohaus, 1910 [4]
        pulchra Arrow, 1911 [3]
        punctata Endrödi, 1968 [1]
      - PYCNOSCHEMA Thomson, 1858 [86]
        andrei Dechambre, 1993 [4]
        antiorii (Gestro, 1881) [4]
        armatum (Thomson, 1859) [1]
        brittoni Endrödi, 1961 [3]
        corydon (Olivier, 1789) [1]
        cuspidatum Thomson, 1859 [3]
        dechambrei Silvestre, 2000 [3]
        desfontainei desfontainei Silvestre, 2000 [1]
        desfontainei ivoiriense Silvestre, 2000 [1]
        diversum cavifrons Lesne, 1906 [3]
        diversum diversum Péringuey, 1901 [4]
        kisoa Dechambre, 1986 [1]
        lacordairei Thomson, 1859 [4]
        mandibularis Dechambre, 1980 [1]
        mashunum Péringuey, 1901 [4]
        mirei Dechambre, 1977 [3]
        mossambicum Prokofiev, 2012 [1]
        operculatum Thomson, 1859 [4]
        parvicornis Fairmaire, 1894 [3]
        politum Arrow, 1906 [4]
        popei Dechambre, 1980 [1]
        prostomis (Fairmaire, 1897) [3]
        rawlinsi Dechambre, 1993 [1]
        schoutedeni Burgeon, 1928 [6]
        scrofa Harold, 1880 [7]
        simplicicolle Kolbe, 1905 [4]
        subulatum Quedenfeldt, 1884 [4]
        suillum Kolbe, 1905 [4]
        uhligi Dechambre, 1993 [1]
        villiersi Dechambre, 1980 [1]
      - SEMANOPTERUS Hope, 1847 [22]
        angustatus Blackburn, 1888 [4]
        calabyi Carne, 1957 [1]
        leai (Blackburn, 1897) [3]
        rectangulus Blackburn, 1895 [4]
        solidus (Burmeister, 1847) [3]
        subcostatus (Castelnau 1840) [3]
        tricostatus Blackburn, 1896 [3]
      - SIRALUS Dechambre, 1997 [1]
        insularis Dechambre, 1998 [1]
      - TEINOGENYS Sharp, 1873 [22]
        annectens Carne, 1957 [1]
        aurilegulus (Blackburn, 1896) [3]
        brevis Sharp, 1873 [1]
        calvus (Blackburn, 1888) [1]
        carinaticeps (Lea, 1919) [1]
        deceptus Carne, 1957 [1]
        demarzi Endrödi, 1971 [1]
        dives (Blackburn, 1896) [1]
        inermicollis (Lea, 1917) [1]
        inermis Blackburn, 1890 [1]
        jenkinsi Carne, 1957 [1]
        laevis (Arrow, 1914) [1]
        metallicola (Blackburn, 1892) [1]
        mooniensis Allsopp & Carne, 1986 [1]
        nitidus Sharp, 1873 [1]
        paradoxus Carne, 1957 [1]
        richardsae (Blackburn, 1890) [3]
      - TEMNORHYNCHUS Hope, 1837 [96]
        acutemarginatus Dechambre, 1976 [1]
        baal baal Reiche & Saulcy, 1856 [4]
        baal sennaariensis Fairmaire, 1868 [4]
        burgeoni Paulian, 1946 [4]
        clypeatus Klug, 1855 [4]
        coquereli Fairmaire, 1868 [3]
        coronatus antiochus Fairmaire, 1868 [1]
        coronatus coronatus (Fabricius, 1781) [4]
        cribratus cribratus Bates, 1884 [4]
        cribratus freyi Endrödi, 1961 [1]
        descarpentriesi Paulian, 1946 [4]
        elongatus Arrow, 1937 [1]
        flavipennis Arrow, 1937 [4]
        grandicornis Fairmaire, 1899 [1]
        kasanganus Krell, 1993 [1]
        luna erectilobus Arrow, 1937 [1]
        luna luna Fairmaire, 1868 [4]
        minor Paulian, 1954 [1]
        overlaeti Burgeon, 1947 [4]
        perrieri Fairmaire, 1899 [3]
        raffrayi Fairmaire, 1887 [3]
        repandus Burmeister, 1847 [3]
        retusus (Fabricius, 1781) [5]
        rugatus Kolbe, 1905 [4]
        scutiger Kolbe, 1905 [3]
        sjöstedti Kolbe, 1910 [4]
        stormsi Duvivier, 1891 [4]
        tridentatus Lansberge, 1886 [4]
        truncatus (Klug, 1832) [3]
        zairensis Krell, 1995 [4]
        zambezianus occidentalis Krell, 1992 [1]
        zambezianus zambezianus Krell, 1992 [3]
      - THRONISTES Burmeister, 1847 [3]
        rouxi Burmeister, 1847 [3]
      - TRISSODON Burmeister, 1847 [23]
        australasiae (Hope, 1837) [4]
        bidentatus Allsopp, 1993 [1]
        bryanti Carne, 1957 [1]
        curtus (Burmeister, 1847) [4]
        glaber (Arrow, 1914) [1]
        howdenorum Allsopp, 1993 [1]
        kiwanus Carne, 1957 [1]
        laevigatus (Burmeister, 1847) [3]
        laticollis (Burmeister, 1847) [1]
        nasutus (Blackburn, 1896) [1]
        setosus Carne, 1957 [1]
        subopacus (Arrow, 1914) [1]
        tibialis Carne, 1957 [1]
        tudor Carne, 1957 [1]
      - WERNORYCTES Dechambre, 1999 [11]
        minettii Silvestre, 2011 [5]
        werneri Dechambre, 1999 [6]
    - PHILEURINI [568]
      - ACTINOBOLUS Westwood, 1841 [14]
        alvarengai Dupuis & Dechambre, 1998 [1]
        barbatus Endrödi, 1978 [3]
        degallieri Dechambre & Lumaret, 1986 [1]
        emarginatus Endrödi, 1978 [1]
        pilosus Dupuis & Dechambre, 1998 [1]
        radians Westwood, 1841 [4]
        spiniformis Dupuis & Dechambre, 1998 [1]
        trilobus Lüderwaldt, 1910 [1]
      - ALLOPHILEURINUS Dupuis & Dechambre, 2001 [2]
        cavifrons Dupuis & Dechambre, 2001 [1]
        mediopunctatus Du & Dechambre, 2001 [1]
      - AMBLYODUS Westwood, 1878 [2]
        castroi Grossi & Grossi, 2011 [1]
        taurus Westwood, 1878 [1]
      - AMBLYOPROCTUS Kolbe, 1910 [30]
        amazonicus Endrödi, 1977 [3]
        arianorum Dechambre, 2008 [4]
        bituberculatus Dechambre, 2008 [1]
        boondocksius Ratcliffe, 1988 [3]
        centroamericanus Ratcliffe, 2003 [1]
        chalumeaui Endrödi, 1977 [3]
        cornutus Dechambre, 2008 [1]
        dechambrei Ponchel, 2009 [1]
        ocellatus Lamant-Voirin, 1995 [1]
        piliger (Perty, 1830) [3]
        rugosus (Erichson, 1847) [3]
        squamosus Dechambre, 2008 [1]
        torulosus Kolbe, 1910 [4]
      - ARCHOPHANES Kolbe, 1905 [4]
        cratericollis (Fairmaire, 1894) [4]
      - ARCHOPHILEURUS Kolbe, 1910 [58]
        alternans Endrödi, 1977 [1]
        aper Endrödi, 1977 [3]
        bifoveatus Endrödi, 1977 [1]
        burmeisteri (Arrow, 1908) [3]
        chaconus (Kolbe, 1910) [7]
        clarionicus Moron, 1990 [1]
        cribrosus (LeConte, 1854) [3]
        elatus (Prell, 1914) [1]
        fimbriatus (Burmeister, 1847) [1]
        fodiens (Kolbe, 1910) [1]
        foveicollis (Burmeister, 1847) [3]
        guyanus Dechambre, 2006 [1]
        kolbeanus (Ohaus, 1910) [1]
        latipennis (Burmeister, 1847) [3]
        oedipus (Prell, 1912) [1]
        opacostriatus (Ohaus, 1911) [3]
        ovis (Burmeister, 1847) [4]
        passaloides (Prell, 1914) [1]
        peruanus Endrödi, 1977 [1]
        petropolitanus (Ohaus, 1910) [1]
        quadrivii Dechambre, 2006 [1]
        simplex (Bates, 1887) [1]
        spinosus Dechambre, 2006 [1]
        sus Dechambre, 2006 [1]
        temnorrhynchodes Lamant-Voirin, 1995 [1]
        tmetoplus (Prell, 1912) [3]
        trituberculatus Endrödi, 1977 [1]
        vervex (Burmeister, 1847) [5]
        wagneri (Ohaus, 1911) [1]
        zirschkai Endrödi, 1981 [2]
      - CAYMANIA Ratcliffe & Cave, 2010 [1]
        nitidissima Ratcliffe & Cave, 2010 [1]
      - CERATOPHILEURUS Ohaus, 1911 [4]
        lemoulti Ohaus, 1911 [4]
      - CHILIPHILEURUS Endrödi, 1977 [1]
        tuberculatus Endrödi, 1977 [1]
      - CNEMIDOPHILEURUS Kolbe, 1910 [1]
        personatus Kolbe, 1910 [1]
      - CRYPTODUS MacLeay, 1819 [46]
        bilobus Lea, 1917 [1]
        caviceps Westwood, 1856 [3]
        concentricus Lea, 1920 [3]
        costulipennis Fairmaire, 1877 [1]
        cycnorum Fairmaire, 1877 [1]
        debilis Blackburn, 1890 [1]
        foveatus Lea, 1919 [4]
        gigas Lea, 1917 [1]
        goerlingi Carne, 1957 [1]
        grossipes Fairmaire, 1877 [1]
        matthewsi Endrödi, 1977 [1]
        oblongoporus Fairmaire, 1877 [1]
        obscurus MacLeay, 1870 [3]
        olivieri (Montrouzier, 1860) [3]
        papuanus Dechambre & Ponchel, 2003 [1]
        paradoxus MacLeay, 1819 [4]
        passaloides Germar, 1848 [3]
        piceus Germar, 1848 [3]
        protensus Fairmaire, 1879 [1]
        relictus Carne, 1957 [1]
        rotundicollis Fairmaire, 1879 [1]
        tasmannianus Westwood, 1841 [3]
        variolosus White, 1841 [3]
      - EOPHILEURUS Arrow, 1908 [93]
        akitai Yamaya & Muramoto, 2008 [1]
        assamensis (Fairmaire, 1898) [1]
        baliensis Yamaya & Muramoto, 2008 [1]
        baolocensis Muramoto, 2004 [1]
        birmanicus Lamant-Voirin, 1995 [1]
        borneensis Endrödi, 1977 [4]
        celebensis Arrow, 1914 [1]
        chinensis chinensis (Faldermann, 1835) [4]
        chinensis irregularis Prell, 1913 [3]
        chinensis okinawanus Nomu. 1964 [4]
        cingalensis cingalensis Arrow, 1908 [4]
        cingalensis decatenatus Arrow, 1910 [1]
        confinis Prell, 1913 [1]
        convexus (Arrow, 1900) [1]
        dentatus (Blackburn, 1895) [3]
        felschei Prell, 1913 [3]
        fenicheli Endrödi, 1977 [1]
        forsteri Endrödi, 1971 [1]
        gracilis Prell, 1913 [1]
        heyrovskyi Kral & Strnad, 1992 [3]
        himalayanus Endrödi, 1977 [1]
        howdeni Yamaya & Muramoto, 2008 [3]
        iwasei Muramoto, 1995 [1]
        javanus Prell, 1913 [4]
        kachinensis Yamaya & Maeda, 2006 [1]
        katsurai Muramoto, 1995 [1]
        malayanus Yamaya & Muramoto, 2008 [1]
        malyi Endrödi, 1978 [4]
        montanus Prell, 1913 [1]
        multidentatus Miyake & Yamaya, 1993 [4]
        nicobarensis Endrödi, 1977 [1]
        niii Yamaya 2008 [1]
        nilgirensis Arrow, 1908 [3]
        oblongus Lamant-Voirin, 1995 [1]
        pectoralis Arrow, 1914 [3]
        perforatus Arrow, 1908 [1]
        planatus (Wiedeman, 1823) [4]
        platypterus (Wiedeman, 1823) [3]
        quadrigeminatus Arrow, 1914 [1]
        ryuheii Muramoto, 2011 [1]
        sondaicus Prell, 1913 [4]
        spatulicornis Lamant-Voirin, 1995 [1]
        sukkiti Yamaya & Maeda, 2006 [1]
        sumbaianus Endrödi, 1977 [1]
        takakuwai Yamaya & Muramoto, 2008 [1]
        tenuiformis Yamaya & Maeda, 2006 [1]
        tetraspermexitus Ratcliffe, 1988 [1]
        thailandensis Endrödi, 1978 [1]
        variolipennis Prell, 1913 [1]
      - GONIOPHILEURUS Kolbe, 1910 [3]
        femoratus (Burmeister, 1847) [3]
      - HAPLOPHILEURUS Kolbe, 1910 [6]
        caudipennis Dupuis, 2011 [1]
        dechambrei Dupuis, 2011 [1]
        uninodis (Burmeister, 1847) [3]
      - HEMIPHILEURUS Kolbe, 1910 [74]
        beckeri (Kolbe, 1910) [1]
        bispinosus Ratcliffe, 2001 [1]
        blandinae Dupuis, 1996 [3]
        brasiliensis Endrödi, 1978 [1]
        caliensis Endrödi, 1985 [1]
        carinatipennis Dupuis & Dechambre, 2000 [1]
        cavei Ratcliffe, 2003 [1]
        cayennensis Endrödi, 1985 [1]
        costatus Endrödi, 1978 [1]
        cribratus (Chevrolat, 1844) [1]
        cubaensis (Chalumeau, 1981) [1]
        curoei Ratcliffe, 2003 [1]
        curvicornis Dupuis & Dechambre, 2000 [1]
        cylindroides (Bates, 1887) [3]
        dechambrei Ratcliffe, 2003 [1]
        dejeani (Bates, 1887) [3]
        depressus (Fabricius, 1801) [3]
        dispar (Kolbe, 1910) [1]
        dyscritus Ratcliffe, 2003 [1]
        elbitae Neita & Ratcliffe, 2010 [1]
        elongatus Dupuis & Dechambre, 2000 [1]
        euniceae Ratcliffe & Cave, 2006 [1]
        gibbosus Dupuis & Dechambre, 2000 [1]
        hiekei Chalumeau, 1988 [4]
        howdeni Endrödi, 1978 [1]
        illatus illatus (LeConte, 1854) [3]
        illatus mexicanus Endrödi, 1978 [1]
        insularis Ratcliffe, 1988 [1]
        isabellae Dupuis, 2004 [1]
        jamaicensis (Howden, 1970) [1]
        kahni Dupuis & Dechambre, 2000 [1]
        laevicauda (Bates, 1887) [1]
        laeviceps (Arrow, 1947) [1]
        laminicornis Dupuis & Dechambre, 2000 [1]
        laticollis (Burmeister, 1847) [4]
        microps (Burmeister, 1847) [1]
        nebulohylaeus Ratcliffe, 2003 [1]
        parvus Dupuis & Dechambre, 2000 [1]
        phratrius Ratcliffe & Ivie, 1998 [1]
        puertoricensis (Chapin, 1935) [1]
        punctatostriatus (Prell, 1914) [1]
        pygidiopunctatissimus Ratcliffe, 2003 [1]
        quadridentatus Ratcliffe, 2001 [1]
        rugulosus Endrödi, 1978 [3]
        ryani Ratcliffe & Ivie, 1998 [1]
        scutellatus Howden, 1978 [1]
        similis Dupuis & Dechambre, 2000 [1]
        simplex (Prell, 1914) [1]
        variolosus (Burmeister, 1847) [4]
        vicarius Prell, 1936 [1]
        vulgatus Dupuis & Dechambre, 2000 [1]
        warneri Ratcliffe, 2001 [1]
      - HOMOPHILEURUS Kolbe, 1910 [24]
        aequatorius Kolbe, 1910 [1]
        amazonicus Kolbe, 1910 [3]
        cubanus Prell, 1911 [1]
        integer (Burmeister, 1847) [3]
        luedeckei Kolbe, 1910 [4]
        luederwaldti (Ohaus, 1910) [1]
        planus Dupuis, 2013 [3]
        quadrituberculatus (Beauvois, 1805) [3]
        tricuspis Prell, 1914 [3]
        waldenfelsi Endrödi, 1978 [1]
      - HOVOPHILEURUS Arrow, 1911 [6]
        hovanus (Fairmaire, 1903) [1]
        latus Dechambre, 1976 [1]
        punctatostriatus Dechambre, 1976 [1]
        strigipennis (Fairmaire, 1905) [1]
        sulcicollis (Fairmaire, 1899) [1]
        sulcipennis Arrow, 1911 [1]
      - MACROCYPHONISTES Ohaus, 1910 [1]
        kolbeanus Ohaus, 1910 [1]
      - METAPHILEURUS Kolbe, 1910 [3]
        explanatus (Burmeister, 1847) [1]
        lacunosus (Burmeister, 1847) [1]
        nitidicollis Kolbe, 1910 [1]
      - MICROPHILEURUS Kolbe, 1910 [2]
        caviceps Kolbe, 1910 [1]
        subulo Prell, 1912 [1]
      - MICTOPHILEURUS Ohaus, 1911 [1]
        punctatus Ohaus, 1911 [1]
      - MORAGUESIA Dechambre, 2008 [4]
        champenoisi Dechambre, 2008 [4]
      - ORYCTOPHILEURUS Kolbe, 1910 [6]
        armicollis Prell, 1911 [1]
        nasicornis (Burmeister, 1847) [3]
        varicosus Prell, 1934 [1]
      - PALAEOPHILEURUS Kolbe, 1910 [17]
        brasiliensis Ratcliffe, 1988 [3]
        carbo Ratcliffe, 2002 [1]
        demezi Dupuis, 2012 [1]
        erebus Ratcliffe, 2002 [1]
        fallax Dechambre, 1996 [4]
        marcusoni Ratcliffe, 1988 [1]
        panamensis Dechambre, 1996 [1]
        proximus Dechambre, 1996 [1]
        sclateri (Bates, 1887) [3]
      - PARAPHILEURUS Endrödi, 1978 [6]
        impressus Endrödi, 1978 [1]
        venezuelensis (Ohaus, 1910) [1]
        ventricosus Endrödi, 1978 [3]
      - PHILEUCOURTUS Dechambre, 2008 [3]
        bicornutus Dechambre, 2008 [3]
      - PHILEURUS Latreille, 1807 [68]
        affinis Burmeister, 1847 [3]
        angustatus Kolbe, 1910 [3]
        bucculentus Ohaus, 1911 [3]
        carinatus carinatus Prell, 1914 [3]
        carinatus declivis Prell, 1914 [4]
        didymus (Linnaeus, 1758) [3]
        endrodii Lamant-Voirin, 1995 [1]
        enigmaticus Dupuis, 2003 [1]
        excavatus Prell, 1911 [1]
        flohri Kolbe, 1910 [1]
        hospes Burmeister, 1847 [3]
        hospitus Prell, 1934 [3]
        incurvatus abscissus Endrödi, 1978 [1]
        kaszabi Endrödi, 1978 [3]
        lecourti Dechambre, 1998 [3]
        limicauda Prell, 1912 [3]
        mundus Prell, 1914 [1]
        patruus Endrödi, 1978 [1]
        rufus Dechambre, 1998 [1]
        schereri Endrödi, 1981 [1]
        toulgoeti Dechambre, 1996 [3]
        truncatus (Beauvois, 1805) [4]
        valgus (Olivier, 1789) [4]
        verus Endrödi, 1978 [4]
        villaemagnae Dechambre, 1998 [3]
        voirinae Endrödi, 1985 [1]
        youngi Ratcliffe, 1988 [4]
        zonianis Ratcliffe, 2011 [1]
      - PLANOPHILEURUS Chapin, 1932 [1]
        planicollis (Chevrolat, 1865) [1]
      - PLATYPHILEURUS Ohaus, 1910 [3]
        felscheanus Ohaus, 1910 [3]
      - PROSPHILEURUS Kolbe, 1905 [3]
        liberianus (Dohrn, 1875) [3]
      - PSEUDOSYRICHTHUS Péringuey, 1901 [13]
        clathratus (Gerstäcker, 1867) [3]
        gerstaeckeri Prell, 1934 [1]
        senegalensis senegalensis (Castelnau, 1840) [7]
        senegalensis variolosus Prell, 1934 [1]
      - RHIZOPLATODES Péringuey, 1901 [9]
        castaneipennis castaneipennis (Boheman, 1857 ) [4]
        castaneipennis foveiceps (Fairmaire, 1893) [3]
        cherlonneixi Dechambre, 1997 [1]
      - RHIZOPLATYS Westwood, 1841 [30]
        arabicus Dechambre, 2005 [4]
        auriculatus (Burmeister, 1847) [4]
        canui Dechambre, 1983 [4]
        cribrarius Westwood, 1841 [4]
        maintyi (Fairmaire, 1898) [3]
        morettoi Silvestre, 1999 [1]
        mucronatus cedrici Dechambre, 1983 [1]
        mucronatus mucronatus (de Beauvois, 1805) [4]
        mucronatus trituberculatus (Burmeister, 1847) [3]
        simplicitarsis (Prell, 1912) [1]
      - SYRICHTHODONTUS Péringuey, 1901 [6]
        cribratus (Boheman,1857) [3]
        lydenburgi Endrödi, 1977 [1]
        spurius (Burmeister, 1847) [1]
      - SYRICHTHOMORPHUS Péringuey, 1901 [3]
        termitophilus Péringuey, 1901 [3]
      - SYRICHTHOSCHEMA Janssens, 1942 [8]
        burgeoni Janssens, 1942 [3]
        katanganum Endrödi, 1977 [4]
      - SYRICTES Prell, 1936 [8]
        curvicornis Endrödi, 1977 [3]
        dentatus Prell, 1936 [1]
        syrichthus (Fabricius, 1775) [3]
      - SYRICTOIDES Endrödi, 1977 [1]
        tarsalis Endrödi, 1977 [1]
      - TRIOPLUS Burmeister, 1847 [3]
        cylindricus (Mannerheim, 1829) [3]
  - Typus Materials [188]
  - Dynastidae ACKNOWLEDGMENTS-REMERCIEMENTS [5]
    - Institut Royal des Sciences Naturelles de Belgique - IRSNB [1]
    - Paul Schoolmeesters [1]
    - Alain Drumont [1]
    - Michel Barré & Stéphane Hanot [1]
    - Martin Hardy [1]
- A Visual Presentation of Euchirinae (Coleoptera, Scarabaeidae) [110]
- The Mammals Of The World [1519]
  - Order Carnivora [750]
    - Suborder Caniformia [335]
      - Family AILURIDAE [18]
        GENUS AILURUS CUVIER, 1825 [18]
        AILURUS FULGENS FULGENS CUVIER, 1825 [18]
      - Family CANIDAE [119]
        GENUS ALOPEX KAUP, 1829 [14]
        ALOPEX LAGOPUS LAGOPUS LINNAEUS, 1758 [14]
        GENUS CANIS LINNAEUS, 1758 [55]
        CANIS LATRANS LESTES MERRIAM, 1897 [7]
        CANIS LATRANS OCHROPUS ESCHSCHOLTZ, 1829 [1]
        CANIS LUPUS ARCTOS POCOCK, 1935 [13]
        CANIS LUPUS LUPUS LINNAEUS 1758 [12]
        CANIS LUPUS OCCIDENTALIS RICHARDSON, 1829 [12]
        CANIS MESOMELAS MESOMELAS SCHREBER, 1775 [6]
        CANIS MESOMELAS SCHMIDTI NOACK, 1897 [4]
        GENUS CHRYSOCYON C.E.H. SMITH, 1839 [9]
        CHRYSOCYON BRACHYURUS ILLIGER, 1815 [9]
        GENUS LYCAON BROOKES, 1827 [8]
        LYCAON PICTUS TEMMINCK, 1820 [8]
        GENUS NYCTEREUTES TEMMINCK, 1839 [8]
        NYCTEREUTES PROCYONOIDES PROCYONOIDES GRAY, 1834 [8]
        GENUS OTOCYON MULLER, 1836 [5]
        OTOCYON MEGALOTIS MEGALOTIS DESMAREST, 1822 [5]
        GENUS SPEOTHOS LUND, 1839 [5]
        SPEOTHOS VENATICUS VENATICUS LUND, 1842 [5]
        GENUS UROCYON BAIRD, 1857 [3]
        UROCYON CINEREOARGENTEUS COSTARICENSIS GOODWIN, 1938 [2]
        UROCYON CINEREOARGENTEUS SCOTTII MEARNS, 1891 [1]
        GENUS VULPES FRISCH, 1775 [12]
        VULPES VELOX SAY, 1823 [8]
        VULPES VULPES CRUCIGERA BECHSTEIN, 1789 [2]
        VULPES ZERDA ZIMMERMANN, 1780 [2]
      - Family MEPHITIDAE [7]
        Subfamily MEPHITINAE [7]
        GENUS MEPHITIS GEOFFROY SAINT-HILAIRE & CUVIER, 1795 [7]
        MEPHITIS MEPHITIS SCHREBER, 1776 [7]
      - Family MUSTELIDAE [48]
        Subfamily LUTRINAE [35]
        GENUS AONYX LESSON, 1827 [9]
        AONYX CINEREUS CINEREUS ILLIGER, 1815 [9]
        GENUS LONTRA GRAY, 1843 [10]
        LONTRA CANADENSIS CANADENSIS SCHREBER, 1776 [10]
        GENUS LUTRA BRISSON, 1762 [9]
        LUTRA LUTRA LINNAEUS, 1758 [9]
        GENUS PTERONURA GRAY, 1837 [7]
        PTERONURA BRASILIENSIS GMELIN, 1788 [7]
        Subfamily MARTINAE [5]
        GENUS GULO PALLAS, 1780 [5]
        GULO GULO GULO LINNAEUS, 1758 [5]
        Subfamily MELLIVORINAE [5]
        GENUS MELLIVORA STORR, 1780 [5]
        MELLIVORA CAPENSIS CAPENSIS SCHREBER, 1776 [5]
        Subfamily MUSTELINAE [3]
        GENUS MUSTELA LINNAEUS 1758 [3]
        MUSTELA ERMINEA INVICTA HALL, 1945 [3]
      - Family PROCYONIDAE [28]
        GENUS NASUA STORR, 1780 [12]
        NASUA NARICA NARICA LINNAEUS, 1766 [5]
        NASUA NASUA NASUA LINNAEUS, 1766 [7]
        GENUS PROCYON STORR, 1780 [16]
        PROCYON LOTOR LOTOR LINNAEUS, 1758 [8]
        PROCYRON LOTOR MEGALODOUS LOWERY, 1943 [3]
        PROCYRON LOTOR PALLIDUS MERRIAM, 1900 [1]
        PROCYON LOTOR SSP [4]
      - Family URSIDAE [115]
        Subfamily AILUROPODINAE [21]
        GENUS AILUROPODA MINE-EDWARDS, 1870 [21]
        AILUROPODA MELANOLEUCA DAVID, 1869 [21]
        Subfamily TREMARCTINAE [16]
        GENUS TREMARCTOS GERVAIS, 1855 [16]
        TREMARCTOS ORNATUS CUVIER, 1825 [16]
        Subfamily URSINAE [78]
        Genus HELARCTOS HORSFIELD, 1825 [6]
        HELARCTOS MALAYANUS RAFFLES, 1822 [6]
        Genus URSUS LINNAEUS, 1758 [72]
        URSUS AMERICANUS ALTIFRONTALIS ELLIOT, 1903 [3]
        URSUS AMERICANUS CINNAMOMUM AUDUBON & BACHMAN, 1854 [14]
        URSUS ARCTOS ARCTOS LINNAEUS, 1758 [11]
        URSUS ARCTOS BERINGIANUS MIDDENDORFF, 1853 [12]
        URSUS ARCTOS HORRIBILIS ORD, 1815 [4]
        URSUS ARCTOS LASIOTUS GRAY, 1867 [12]
        URSUS MARITIMUS PHIPPS, 1774 [16]
    - Suborder Feliformia [414]
  - Hoofed Mammals [769]
- The Genus Morpho [127]
- Ecureuils Du Monde [136]
  - Les Ecureuils Du Monde [136]
- Oiseaux Du Monde [426]
  - Oiseaux Des Etats-Unis [90]
  - Oiseaux De Tanzanie [50]
  - Oiseaux Du Sri Lanka [25]
  - Oiseaux Du Salvador [8]
  - Oiseaux Du Panama [4]
  - Oiseaux Du Nicaragua [5]
  - Oiseaux Du Laos [4]
  - Oiseaux Du Kenya [57]
  - Oiseaux Du Honduras [6]
  - Oiseaux Du Guatemala [28]
  - Oiseaux Du Costa Rica [30]
  - Oiseaux Du Canada [25]
  - Oiseaux Du Cambodge [1]
  - Oiseaux Du Botswana [19]
  - Oiseaux De Polynésie [3]
  - Oiseaux De Namibie [51]
  - Oiseaux De Martinique [11]
  - Oiseaux De Madère [3]
  - Oiseaux De Bali [5]
  - Livre d'Or [1]
Nos voyages au bout du monde [14901]
- Afrique du Sud - 2003 [1]
- Australie - 2018 [1]
- Autriche - 1997 - 1998 [1]
  - TYROL [1]
- Bali - 2012 [556]
  - Lovina [11]
  - Sanur [12]
  - Amed [6]
  - Ubud [2]
  - Village fortifié de Tenganan [15]
  - Temples et Palais [166]
    - Temple familial [12]
    - Brahmavihara Arama [16]
    - Yeh Pulu [6]
    - Pura Ulun Danu Bratan [6]
    - Taman Tirtagangga [17]
    - Pura Dalem Segara Madhu [19]
    - Pura Gunung Kawi [9]
    - Puri Saren [7]
    - Pura Tirta Empul [24]
    - Pura Taman Saraswati [2]
    - Pura Segara [11]
    - Pura Luhur Muncaksari [8]
    - Pura Blanjong [11]
    - Pura Taman Ayun [15]
    - Pura Luhur Batukau [3]
  - Folklore et tradition [100]
    - Cérémonie de l' Odolan [26]
    - La danse du Barong [28]
    - Cérémonie du limage des dents [10]
    - Mariage balinais [18]
    - Combat de coqs [9]
    - Maison balinaise [9]
  - L' artisanat [15]
  - Récolte de Sel [8]
  - La faune [9]