Un Monde En Couleurs

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Lucanus (Lucanus) Scopoli, 1763

 

  • adelmae Zilioli, 2003

    4 photos

     

    Lucanus adelmae

     

     

    Lucanus adelmae , ZILIOLI M. 2003.

    Ø       Myanmar

                                                                                                                                                

    Bibliography :

     

    ZILIOLI M. ( 2003 ) A description of Lucanus adelmae n. sp., a new stag-beetle from Myanmar (Coleoptera Lucanidae). Atti Soc. nat. Museo civ. Stor. nat. Milano, 2003; 144(1): 83-89, 6 figg.

     

    GEKKAN MUSHI ( 2003 ) Suite a The Lucanid beetles of the world.de MIZUNUMA T. & NAGAI S. Gekkan Mushi, 390: 2-17

     

    HUANG H. ( 2006 ) New descriptions and notes on Chinese stag-beetles, with discovery of the second species of Noseolucanus from SE Tibet. Coleoptera, 10(2006): 11-34, 19 figg.

     

    TEXT : GALANT ALAIN 

     

    PICTURES : GALANT ALAIN

     

  • angusticornis ssp. angusticornis Didier, 1925

    7 photos

     

    Lucanus (Lucanus) angusticornis

     

     

    Lucanus (Lucanus) angusticornis DIDIER, 1925.

    Lucanus angusticornis (Boileau in litt.) DIDIER, 1925:219 [Laos : Xieng-Khouang].

    Lucanus angusticornis DIDIER; Didier, 1926:144

    Lucanus angusticornis DIDIER; Didier & Séguy, 1953:81

    Lucanus angusticornis DIDIER; Benesh, 1960:131

    Lucanus angusticornis DIDIER; Lacroix, 1972:50

    Lucanus angusticornis DIDIER; Kurosawa, 1978:143

    Lucanus angusticornis DIDIER; Sakaino, 1981:23

    Lucanus angusticornis DIDIER; Maes, 1992:15

    Lucanus angusticornis DIDIER; Kawano, 1994:9

    Lucanus angusticornis DIDIER; Mizunuma & Nagai, 1994:216, pl.15

    Lucanus angusticornis DIDIER; Kawano, 1997:454

    Lucanus angusticornis DIDIER; Kawano, 1997:25.

    Lucanus angusticornis DIDIER; Araya, 2001:10, figs.1-5.

    •Laos : Xieng-Khouang, 18-VI-1919, ex. coll. Boileau (male HOLOTYPE of Lucanus angusticornis [Boileau in litteris] DIDIER, 1925 en Museum National d'Histoire Naturelle, Paris).
    •Vietnam : Tonkin : Tam Dao near Vinh Yen at NE Hanoi, leg. H. Perrot (female TYPE of Lucanus angusticornis DIDIER, 1925 by LACROIX, 1969, published in 1972 en Museum National d'Histoire Naturelle, Paris).

    Bibliography :

    ARAYA K. (2001) Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae) from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.

     

    TEXT : MAES JEAN-MICHEL

     

    PICTURES : GALANT ALAIN

     

  • angusticornis ssp. inclinatus Schenk, 2008

    5 photos

    Lucanus angusticornis inclinatus Schenk, 2008

     

    China, Yunnan.

     

    SCHENK K.D., 2008. Beitrag zur Kenntnis der Hirschkäfer Asiens und Beschreibung mehrerer neuer Arten; Beetles World 1: 1-12.

     

     

    TEXT : GALANT ALAIN 

     

    PICTURES : GALANT ALAIN

  • brivioi ssp. brivoi Zilioli, 2003

    4 photos

     

    Lucanus (Lucanus) 

     

     

    Lucanus brivioi , ZILIOLI M. 2003.

    Ø       China, Fujian

     

    Bibliography :

     

    ZILIOLI M. ( 2003 )  Lucanus brivioi n.sp., a new stag beetle from the Wuyi shan mountains, Fujian, China ( Coleoptera Lucanidae ). Atti Soc. it. Sci. nat. Museo civ. Stor. nat. Milano, 144 (II): 265-272, Dicembre 2003.

     

     

     

    TEXT : GALANT ALAIN

     

    PICTURES : GALANT ALAIN

     

  • cantori ssp. cantori Hope, 1842

    5 photos

     

    Lucanus (Lucanus) cantori cantori

     

     

    Lucanus (Lucanus) cantori ssp. cantori HOPE, 1842 [Khasi Hills].

    Lucanus cantori HOPE, 1842:83 [Khasi Hills].

    Lucanus cantori HOPE; Hope, 1843:363

    Lucanus cantori HOPE; Hope, 1845:73

    Lucanus cantori HOPE; Hope & Westwood, 1845:5

    Lucanus cantori HOPE; Burmeister, 1847:527

    Lucanus cantori HOPE; Planet, 1899:275

    Lucanus cantori HOPE; Planet, 1902:57, pl.13

    Lucanus cantori HOPE; Heyne & Taschenberg, 1908:50, pl.6

    Lucanus cantori HOPE; Gravely, 1915:411

    Lucanus cantori HOPE; Didier, 1949:pl.12

    Lucanus cantori HOPE; Arrow, 1950:51, pl.3

    Lucanus cantori HOPE; Benesh, 1960:131

    Lucanus cantori HOPE; Lacroix, 1967:97-98

    Lucanus cantori HOPE; Maes, 1992:15-16

    Lucanus cantori HOPE; Mizunuma & Nagai, 1994:213, pl.10.

    Lucanus cantori HOPE; Araya, 2001:11, fig.13.

    •India : West Bengal : Darjeeling
    •India : West Bengal : Kurseong
    •India : West Bengal : Gopaldhara
    •India : West Bengal : Rungbong Valley
    •India : West Bengal : Darjeeling : Redong
    •India : "?Himalaya"or "Khasi Hills" (male TYPE of Lucanus cantori HOPE, 1842, # 356 in Oxford University Museum of Natural History).
    •Bhutan : Pedong
    •Bhutan : Paro Distr.
    •Bhutan : Gedu.

    Adults on ? : Castanopsis, Symplocos.

    Bibliography :

    ARAYA K. (2001) Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae) from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.

     

    TEXT : MAES JEAN-MICHEL

     

    PICTURES : GALANT ALAIN

     

  • cantori ssp. colasi Lacroix, 1967

    4 photos

     

    Lucanus (Lucanus) cantori colasi

     

     

    Lucanus (Lucanus) cantori ssp. colasi LACROIX, 1967.

    Lucanus cantori ssp. colasi LACROIX, 1967:97-98 [India : Shillong].

    India : Meghalaya : Shillong (male TYPE of Lucanus cantori ssp. colasi LACROIX, 1967).
    India : Meghalaya : Khasi Hills.

    Note : it seems that the nominal subspecies was describe from Khasi Hills, if it is so, colasi will be synonym of cantori and then the ssp. of Darjeeling is unnamed.

     

    TEXT : MAES JEAN-MICHEL

     

    PICTURES : GALANT ALAIN

     

  • cervus f. scapulodonta Weinreich, 1963

    5 photos

     

    Lucanus ( Lucanus ) cervus f. scapulodonta 

     

     

    Lucanus cervus f. scapulodonta WEINREICH, 1963:29.

    Burton, 1965:121; Clark, 1966:199

    Lucanus cervus LINNAEUS; Bahuena, 1967

    Clark, 1967:24; Lacroix, 1967:98

    Lucanus cervus (LINNAEUS); Freude, 1968:59-64

    Lacroix, 1968:234-235; Lacroix, 1969:119-129; Machatschke, 1969; Mathieu, 1969:1061; Freude, 1971:283

    Lucanus cervus (LINNAEUS); Cofais, 1972:32-33

    Lovari, 1972:36

    Lucanus cervus (LINNAEUS); Mal, 1972:161-167

    Lacroix & Bomans, 1973:152; Kersmaekers, 1973:219; Verstraeten, Boosten & Gaspar, 1973:mapa 713; Jacobs & Renner, 1974; Radisic, 1974; Harde, 1975:16; Athias, 1976:194; Nadolski, 1976; Andersson, 1977:100

     

    TEXT : MAES JEAN-MICHEL

     

    PICTURES : GALANT ALAIN

     

  • cervus ssp. akbesianus Planet, 1896

    3 photos

     

    Lucanus (Lucanus) 

    cervus akbesianus

     

     

    Lucanus (Lucanus) cervus ssp. akbesianus PLANET, 1896.

    Lucanus cervus turcicus ab. syriacus MOTSCHULSKY (nec MOTSCHULSKY) 1897.

    Lucanus cervus ab. motschulskyi BENESH 1960.

    Lucanus poujadei PLANET 1897 [Syrie].

    Lucanus smyrniacus DIDIER & SEGUY 1953 [Asie mineure].

    Lebanon : Djezin;

    Syria : Akbes* = Sgée, Marah, Smyrne, Alexandrette, Ghazir, Marsch.

     

    TEXT : MAES JEAN-MICHEL 

     

    PICTURES : GALANT ALAIN

     

  • cervus ssp. cervus (Linnaeus, 1758)

    9 photos

     

    Lucanus (Lucanus) cervus cervus

     

     

    Lucanus (Lucanus) cervus ssp. cervus (LINNAEUS, 1758).

    Scarabaeus cornutus DALE, 1693:389, pl.17.

    Linnaeus, 1745:69; Rossel, 1746:25, pl.4-5

    Scarabaeus cervus LINNAEUS, 1758:353.

    Poda, 1761:21; Sulzer, 1761:3, pl.1; Geoffroy, 1762:61, pl.1; Pontopp., 1763:430; Kurz., 1765:199; Scopoli, 1763:1; De Bomare, 1764:486; Gronov, 1764:147; Muller, 1764:2; Schaeffer, 1766:pl.9; Linnaeus, 1767:559; Schluga, 1767:pl.1; De Geer, 1774:327, pl.12; Fabricius, 1775:1

    Lucanus capreolus FUESSLY, 1775:3

    Lucanus dorcas MULLER, 1776:52

    Sulzer, 1776:19, pl.2; Bergstrasser, 1778:18, 25, pl.4, 13; Fuessly, 1778:63; Schaeffer, (1766)-1779:pl.133; Barbut, 1781:15, pl.2; Fabricius, 1781:1; Fuessly, 1781:2; Schrank, 1781:32; Von Laichart., 1781:1; Von Scheven, 1782:60; Panzer, 1783:36, pl.29

    Lucanus cervus LINNAEUS; De Asso, 1784

    Leske, 1784:464; Bonsd., 1785:221; Fourcroy, 1785:2; Von Moll, 1785:149; Fabricius, 1787:1; Petagna, 1787:1; Von Meidinger, 1787:117; Zschach, 1788:1; De Villers, 1789:41

    Lucanus capra OLIVIER, 1789:9-12, pl.1-2

    Schrank, 1789:374; Von Razoum., 1789:138; Brahm, 1790:124, 130

    Lucanus armiger HERBST, 1790:287, 289, 299, 301, pl.D-E, pl.33-34.

    Lucanus hircus HERBST, 1790:287, 289, 299, 301, pl.D-E, pl.33-34.

    Linnaeus (ed. Gmelin), 1790:1588; Preyssler, 1790:9; Roesel, 1790:1749, pl.iv; Rossi, 1790:1-2; Scriba, 1790:40; Mayer, 1791:181; Donovan, 1792:31, pl.13; Fabricius, 1792:236-237; Kugelann, 1792:255; Martyn, 1792:pl.5; Brahm, 1793:657; Fabricius, 1793:236; Schwarz, 1793:84; Panzer, 1795:244; Hoppe, 1796:169, 179; Hoppe, 1797:179, 200; Panzer, 1798:11;

    Olivier (ed. Illiger), 1800:55-59; Paykull, 1800:45; Walther, 1800:352; Fabricius, 1801:248-249

    Lucanus inermis MARSHAM, 1802:46, 48, pl.2 [Loc. ?].

    Blumenb., 1803:321; Latreille, 1804:246, pl.86; Posselt, 1804:15, pl.ii; Duftschm., 1805:64; Panzer, 1805:110; Bechst. & Scharfenb., 1805:80; Sturm, 1805:pl.1-2; Thunberg, 1806:190-191; Fallen, 1807:12, 22

    Lucanus grandis HAWORTH, 1807:77

    Latreille, 1807:135; Gyllenhall, 1808:65-66; Schonherr, 1817:318, 320-321; W.S. Macleay, 1819:114; Blot, 1824:84; Muller, 1825:553-672; Klug, 1829:365, pl.15; Stephens, 1830:166; Davis, 1832:86; Curtis, 1834:490; Audoin, 1835:81; Pictet, 1836:393; Dejean, 1837:193; Mulsant, 1839:119, pl.2; Newport, 1839:853-994; Ratzeburg, 1839:105; Westwood, 1839:187; Castelnau, 1840:171; Spry & Shuckard, 1840:34, pl.42; Heer, (1838)-1842:495

    Lucanus microcephalus MULSANT, 1842:8, 583, 586, pl.3

    La Ferté Senectère en Mulsant, 1842:8; Burmeister, (1835)-1843:pl.24; Sturm, 1843:347, pl.5; Tulk, 1843:451; Brulle, 1844:271-374; Desmarest, 1845:53

    Lucanus americanus HOPE & WESTWOOD, 1845:9-10 [N. America, error]

    Lucanus lusitanicus HOPE & WESTWOOD, 1845:9-10 [Portugal].

    Lucanus vicinus HOPE & WESTWOOD, 1845:9-10 [Poonah, error].

    Motschulsky, 1845:60; Bellier de la Chavignerie, 1846:28; Blanchard, 1846:322

    Lucanus major BURMEISTER, 1847:350, 527

    Erichson, 1848:936, 938; Kriechbaun., 1848:208;

    Reiche, 1853:71; Perris, 1854:105; Mulsant & Godart, 1855:250; Mulsant & Godart, 1855:150; Lacordaire, 1856:23; Reiche, 1856:81; Delarouzee, 1859:98; Delarouzee, 1859:42; Duval, 1859:7, 9-10, pl.1; Gerst., 1859:110; Deyrolle, 1860:22; Kraatz, 1860:68, 265, 273, pl.7; Parry, 1864:71; Abeille de Perrin, 1867:65, 70; Cornelius, 1867:435; Cornelius, 1868:33; Landois, 1868:88-95, pl.vi; MacLachlan, 1868:ix; Motschulsky, 1870:25-31, 34, pl.2; De Narcillac, 1871:186; Hochuth, 1871:85; Mulsant & Rey, 1871:8, 14, 16; Schioedte, 1873:342-345, pl.15; Redtenbacher, 1874:451; Schiodte, 1874:341, pl.xviii-xix; Schlechtendal, 1874:25; Schlechtendal, 1875:158; Kraatz, 1876:72; Rossi, 1876:95; Schmidt-Goebel, 1876:392; Perris, 1877:101-102, pl.5; Rossi, 1878:227; Schreiber, 1878:259; Brandt, 1879:1-17; Cotton, 1880:83-84; Reichenau, 1880:55; Rupertsberger, 1880:142; Hammond, 1881:14-26, pl.2; Reichenau, 1881:172-194; Kraatz, 1883:27; Leboeuf, 1883:45; Strubing, 1883:159; Cornelius, 1884:33; Smith, 1885:27; Frivaldsky, 1886:78; Gadeau de Kerville, 1886:176; Fallou, 1889:141; Rousseau, 1889:170; Fowler, 1890:4; Planet, 1890:156-157, 164-165, 215-216

    Lucanus antennatus REY, 1890:164

    Reitter, 1891:142-143; Verhoeff, 1893:113-170; Verhoeff, 1893:209-260; Binet, 1894:449-580; Pottier, 1894:27; Rupertsberger, 1894:146; Borgschulze, 1895:131; Caradja, 1895:77; Eggers, 1895:131; Schaufuss, 1895:111; Selmons, 1895:52; Planet, 1896:11, 99, 168, 238, 256; Planet, 1897:64, 192, 229, 267; Weise, 1897:389; Bothe, 1899:140; Planet, 1899:31, 35, 42, 46, 55, 58, 62, 64-65, 104, 107, pl.9, 10, 14; Rupertsberger, 1899:235; Stichel, 1899:19

    Lucanus longipennis VON ROTHENBURG, 1900:99

    Jarder, 1901:28; Le Senechal, 1901:190; Xambeau, 1901:604; Planet, 1902:52, 100, 111, 114-115, 119, 120, 125; Weber, 1902:335; Biedermann, 1903:395-481

    Lucanus cervus LINNAEUS; Champion, 1903

    Lucanus cervus LINNAEUS; Champion, 1904

    Marchand, 1904:37; Rudow, 1904:179, 187, 196; Schuster, 1904:338; Amman, 1905:139; Scabra, 1905:7-8, pl.; Rey, 1906:59; Barthe, 1909:5-6; Chagrin, 1909:404; Reitter, 1909:299, pl.68; Chinaglia, 1910:1-8, 25-39; Leoni, 1910:153

    Lucanus capitalis MOLLENKAMP, 1910:242, 575

    Schieser, 1910:29; Bedel, 1911:8; Kapzov, 1911:297-337; Krizenecky, 1911:113; Pic, 1911:399; Sasse, 1911:69-104

    Lucanus validus MOLLENKAMP, 1912:302 [Borneo, error].

    Muzik, 1912:189; Ramme, 1912:37; Roubal, 1912:386; Sirguey, 1912:134; Boileau, 1913:218, 220; Bordas, 1913:358-364; Fairmaire & Planet, 1913:195, pl.9; Kuhnt, 1913:380; Bordas, 1914:181-183; Braatz, (1913)-1914:267; Moritz, 1914:138; Stellwag, 1914:359-429; Chinaglia, 1915:7; Houlbert & Monnot, 1915:4, 6; Labitte, 1916:105; Schaufuss, 1916:1252, pl.48; Verhoeff, 1916:354-372; Della Beffa, 1921:3; Houlbert (1912)-1921:167; Dudich, 1923:129; Grutzner, 1924:130; Nagel, 1924:398-399; Henriksen, 1925:134-143; Nagel, 1928:273

    Lucanus ornatus BERNAU, 1929:72

    Catherine, 1929:5-6; Vignon, 1929:89-125; Gadeau de Kerville, 1930:63; Nagel, 1930:90; Goldschmidt, 1931

    Lucanus nigrinus MARCH, 1931.

    Marcu, 1931:214; Portevin, 1931:2-3, pl.6; Roubal, 1931:36; Baudrimont, 1932:116; Franz, 1932:134; Joy, 1932:242; Joy, 1932:pl.69; Kraemer, 1932:321; Kraemer, 1932:86-117; Porta, 1932:371; Didier, 1933:24, pl.1; Hepp, 1933:126-127; Hepp, 1934:84; Golovyanko, 1935:38; Van Emden, 1935:194, 200; Hepp, 1936:173; Paulian, 1936:751-759; Didier, 1937:xxxviii, xlii; Muller & Schoop, 1937:41-116; Palm, 1941:17; Van Emden, 1941:119; Garnier, 1945:110; Sellier & Razet, 1947:223-225; Colas, 1949:128-131; Didier, 1949:pl.1, 3;

    Leiler, 1950:157; Muller & Schoop, 1950:17-18; Arrow, 1951:pl.8; Condrillier, 1951:20; Brull, 1952; Didier & Séguy, 1952:pl.15, 17, 24; Tressens, 1952:57; Auber, 1953:99-100; Didier & Séguy, 1953:27-32; Tippmann, 1954; Theodorides, 1955; Britton, 1956:6; Landin, 1956:1; Paulian, 1956:36; Freude, 1957:129-134; Harz, 1957:22-23; Dorn, 1958:62-64

    Lucanus cervus LINNAEUS; Español, 1958

    Lucanus cervus (LINNAEUS); Horion, 1958

    Papperitz, 1958; Weise, 1958; De Ligondes, 1959:52-56; Franz, 1959:74-79; Marie, 1959:125-127; Paulian, 1959:38-39; Weinreich, 1959:282-283; Benesh, 1960:132, 138-140; Janssens, 1960:95; Muspratt, 1960:53; Kuhn, 1961; Colas, 1962:118; Demoulin, 1963:pl.8

    Lucanus cervus f. scapulodonta WEINREICH, 1963:29.

    Burton, 1965:121; Clark, 1966:199

    Lucanus cervus LINNAEUS; Bahuena, 1967

    Clark, 1967:24; Lacroix, 1967:98

    Lucanus cervus (LINNAEUS); Freude, 1968:59-64

    Lacroix, 1968:234-235; Lacroix, 1969:119-129; Machatschke, 1969; Mathieu, 1969:1061; Freude, 1971:283

    Lucanus cervus (LINNAEUS); Cofais, 1972:32-33

    Lovari, 1972:36

    Lucanus cervus (LINNAEUS); Mal, 1972:161-167

    Lacroix & Bomans, 1973:152; Kersmaekers, 1973:219; Verstraeten, Boosten & Gaspar, 1973:mapa 713; Jacobs & Renner, 1974; Radisic, 1974; Harde, 1975:16; Athias, 1976:194; Nadolski, 1976; Andersson, 1977:100

    Lucanus cervus (LINNAEUS); Clark, 1977:9

    Leech, 1977:69; Perrier, 1977:167; Hempel & Schiemenz, 1978; Lacroix, 1978:253; Cameron, 1980:31; De Walsche, 1980:201; Krikken, 1980:24

    Lucanus cervus (LINNAEUS); Roer, 1980:248-251

    Numhauser, 1981:74; Boosten, 1982:96; Decelle, 1982:203; Español & Belles, 1982:73; Klausnitzer, 1982:14, 17; Bessonnat, 1984:77-80; Lojtnant, 1984; Fellenberg, 1985:79-80; Stolk, 1985:159; Bartolozzi, 1986:12; Cruysbergh, 1986:103; Chalmers-Hunt, 1987:184; Creutz, 1987:58-59; Long, 1987:323-324; Marples, 1987:149-150; Newton, 1987:371; Sakaino, 1987:16; Alexay, 1988:24-25; Andersson, Coulianos, Ehnstrom, Hammarsted, Imby, Janzon, Lindelow & Walden, 1987:70; Gjurasin, 1988:60-61; Grayson, 1988:224; Rabil, 1988:217; Verdcourt, 1988:36; Wootton, 1988:113; Bartolozzi, 1989:104; Carriere, 1989:19-20; George, 1989:28; Pfaff, 1989:31-32; Baker, 1990:12; Hein, 1990:34; Verdcourt, 1990:196

    Lucanus cervus (LINNAEUS); Zemp, Hartmann, Knecht & Sprecher, 1990.

    Bowdrey, 1991:300; Gouranton, Folliot & Thomas, 1991:102-109; Kranjcev, 1991:18-19; Kuhnel, 1991:209; Mamonov, 1991:157-163; Morris, 1991:106; Newton, 1991:36-38; Shepard & Biggs, 1991:7-8; Delatour, 1992:75; Alvarez-Lao, 1992:316-323; Maes, 1992:16-17; Spooner, 1992:329-330; Baraud, 1993:61; Bartolozzi & Onore, 1993:126

    Lucanus cervus (LINNAEUS); Ochse, 1993:47-52.

    Whitehead, 1993:206; Wuest, 1993:132; Hecq, 1994:455; Karner, 1994:71-72; Mizunuma & Nagai, 1994:212, pl. 8-9

    Lucanus cervus (LINNAEUS); Klausnitzer, 1995.

    Viejo & Sanchez Cumplido, 1995:186; Bartolozzi, 1996:486; Teunissen & Smit, 1996:134-135; Araya, 1996:19; Köteles & Bokony, 1996:99; Kral & Vitner, 1996:425

    Lucanus cervus (LINNAEUS); López-Colón, González Peña & Beltrán Valen, 1996:16, 18.

    Lucanus cervus cervus (LINNAEUS); López-Colón, González Peña & Beltrán Valen, 1996: 17.

    Cecchi & Bartolozzi, 1997:122; Heynderycx, 1997:392-393, 396; Hoekstra, 1997:93-95.

    Lucanus cervus LINNAEUS; Sprecher-Uebersax & Durrer, 1998:142-166.

    Lucanus cervus LINNAEUS; Sprecher-Uebersax & Durrer, 1998:471-479.

    Lucanus cervus LINNAEUS; Sprecher-Uebersax & Durrer, 2001:2-11.

    Lucanus cervus LINNAEUS; Sprecher-Uebersax & Durrer, 2001:161-182.

    Lucanus cervus (LINNAEUS); Reyes-Castillo, 2002:486.

    Lucanus cervus (LINNAEUS); Ballerio, 2003:68, 73, 82.

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    Denmark

    Great Britain : Borders (Eccles--55°37N-2°22W), Northumberland (Horsley--54°59N-1°09W), Cumbria (Keswick--54°36N-3°07W), W. Yorks (Burley--53°55N-1°41W, Apperley--53°51N-1°42W), Lancashire (Thornton--53°53N-3°01W), Clwyd (Prestatyn--53°20N-3°23W), Lincoln (Lincolnshire--53°13N-0°31W, Boston--52°59N-0°00--Billups), Nottingham (Nottingham--52°58N-1°09W), Derby (Derbyshire--52°56N-1°28W, Calke, Calke--Garneys), Shropshire (Wellington--52°40N-2°30W {Ercall wood, Apley wood, Ollerton wood}, Church Stretton--52°33N-2°49W), Hereford & Worcester (Bewdley--52°22N-2°20W, Bewdley--52°22N-2°20W--Blatch, Worcestershire--52°11N-2°14W), Warwick (Warwick--52°18N-1°34W, Warwick--52°18N-1°34W--Martineau), Cambridge (Cambridgeshire--52°12N-0°08E--Bloomfield), Buckingham (Buckinghamshire--52°01N-1°00W), Suffolk (Woodbridge--52°06N-1°18E, Ipswich--52°04N-1°09E, Nayland--51°58N-0°52E, Felixstowe--51°58N-1°20E), Essex (Halstead--51°57N-0°21E, Manningtree--51°56N-1°04E, Colchester--51°54N-0°54E, Braintree--51°53N-0°33E, Coggeshall--51°52N-0°41E, Marks Tey--51°52N-0°48E, Brightlingsea--51°49N-1°01E, Witham--51°48N-0°39E, West Mersea--51°47N-0°55E, Epping Forest--51°42N-0°06E, Brentwood--51°37N-0°18E, Tilbury--51°27N-0°22E), Gloucester (Cheltenham--51°54N-205°W, Birdlip--51°49N-2°07W, Newnham--51°48N-2°28W, Hardwicke--51°48N-2°18W), Hertford (Hatfield--51°47N-0°14W, Waltham Cross--51°42N-0°11W), Oxford (Oxford--51°46N-1°15W, Wallingford--51°36N-1°09W, Mapledurham Park), London (Enfield--           51°39N-0°05W, Pinner--51°36N-0°24W, Harrow--51°35N-0°21W = Harrow North, Ilford--51°34N-0°08E, Romford--51°34N-0°11E, Uxbridge--51°33N-0°29W, Upminster--51°33N-0°15E, London--51°32N-0°15W, Southall--51°31N-0°23W, Dagenham--51°31N-0°01E, Woolwich--51°29N-0°03E, Greenwich--51°28N-0°01E, Eltham--51°27N-0°03E, Hounslow--51°25N-0°21W, Hampton--51°25N-0°21W = Hampton Court, Kingston--51°25N-0°18W, Streatham--51°25N-0°08W, Bromley--51°25N-0°01E, Sidcup--51°25N-0°07E,       Wimbledon--51°23N-0°12W, Croydon--51°22N-0°06W, Sutton--51°21N-0°11W, Purley--51°20N-0°07W, Hatton Garden, Cadogan Square, New Forest), Glamorgan (Swansea--51°37N-3°56W), Buckingham (Marlow--51°35N-0°46W), Berkshire (Cookham--51°34N-0°42W, Maidenhead--51°31N-0°43W {Kimbers--Spring Hill}, Slough--            51°31N-0°36W, Pangbourne--51°29N-1°07W {Tidmarsh}, Windsor Forest--51°29N-0°37W, Datchet--51°29N-0°35W, Caversham--        51°28N-0°59W = Caversham Warren, Reading--51°27N-0°58W {Tilehurst}), Kent (Dartford--51°26N-0°14E, Gravesend--51°26N-0°22E, Rochester--51°25N-0°35E, Sheppey I.--51°25N-0°50E {Sheerness--51°26N-0°46E}), Surrey (Staines--51°26N-0°29W, Chertsey--51°24N-0°29W, Surbiton--51°24N-0°19W, Weybridge--51°23N-0°35W, Walton--51°23N-0°35W, Bagshot--51°22N-0°42W = Bagshot Heat, Epsom--51°20N-0°17W, Woking--51°19N-0°25W, Leatherhead--51°18N-0°20W, Box Hill--51°16N-0°20W, Merrow--51°15N-0°32W, Godalming--51°12N-0°37W, Chiddingford--51°07N-0°39W, Merton Park, Kew, Great Leight Rectory), Hampshire (Farnborough--51°17N-0°46W, Winchester--51°05N-1°19W, Bitterne--50°55N-1°22W, Southampton--50°55N-1°24W, Havant--50°52N-0°59W, New Forest--50°51N-1°35W, Beaulieu Abbey--50°49N-1°28W, Gosport--50°48N-1°08W, Lymington--50°46N-1°33W, Southsea--50°46N-1°04W), Somerset (Castle Cary--51°06N-2°31W), Wilts (Salisbury--51°05N-1°48W), Devon (Instow--51°03N-4°10W, Merton--50°53N-4°06W = Merton Park), W. Sussex (Midhurst--50°59N-0°45W, Arundel--50°51N-0°35W, Chichester--50°50N-0°48W, Shoreham-by-See--50°50N-0°17W = Shoreham, Worthing--50°49N-0°22W, West Tarring), E. Sussex (Brighton--50°46N-0°08W), Dorset (Lyme Regis--50°43N-2°57W), Cornwall (Yelverton--50°29N-4°05W, Saltash--50°24N-4°12W, Polperro--50°19N-4°31W, Truro--50°14N-5°03W), Middlesex (Teddington--Fulwell Vicarage, Heston, Lampton), Redmarley, Little Tring, Puttenham, Appleslade, Ellingham, Highwood, Linwood, Calmore, Hamble, Brightwell, Purbrook, Baughurst, Tadley, Mapledurham, Tidmarsh, Ewelme, Swyncombe Down, Bray, Henley, Eashing, Frimley, Wellington College, Brookwood, Sunninghill, Stoke Podges, Buryan, Fulwell, Mount Edgcumb, Bere Alston, Buckland Monachorum, Fingle Gorge, Slapton, Colyton, Parkestone, West Howe, Mudeford, Bransgore, West Cowes, Alverstoke, Kirby Bedon, Broxbourne, Chesnut, Wormley, Great Leighs, West Stow, Layer Breton, Layer-de-La-haye, Peldon, Tollesbury, Stanway, Boxted, Great Horkesley, Mount Bures, East Mersea, Fingringhoe, Arleigh, Elmstead, East Donyland, Thorpe-le-Soken, Wivenhoe, Dedham, Lawford, Thorrington Street, Kersey, Great Bentley, Little Bentley, Bradfield, Brantham, Holbrook, Woolverstone, Little Oakley, Guilford, Wisley, Oatlands, Shepperton, Northwood, Lancing, Sompting, Bramber, Henfield, Steyning, Brockham, Ashtead, Bookham, Ranmore, Chessington, Claygate, East Molesley, Esther, Hanworth, New Malden, Teddington, Fetcham, Heston, Kew, Lampton, Petersham, Richmond Park, Sheen, Carshalton, Mitcham, Barnes, Battersea, Cadogan Square, Chelsea, Chiswick, Fulham, Putney, Roehampton Lane, Wandsworth, Wandsworth Common, Addiscombe, Beckenham, Beddington, Norbury, Stoneleigh, Blackheat, West Dulwich, Bostall Heat, Brockley, Catford, Camberwell, Lambeth, Lee, Lewisham, South Norwood, Southwark, Sydenham, Hatton Garden, Holborn, Chingford, Barcombe, Abbey Wood, Charlton, Chislehurst, Danson Park, Plumstead Common, Upper Belvedere, Leyton, Loughton, Theydon Bois, Lullingstone, Darenth Park, Harold Wood, Larkfield, Grays, Stifford, West Thurrock, Aylesford, Birling, Ditton;

    Netherlands : Drenthe, Overijssel--Twente, Gelderse Vallei, Utrecht (Utrechtse Heuvelrug), Noord-Brabant, Zuid-Limburg, Eysden, Alpesdroom;

    Belgium : Anvers (Merxem, Postel), Limbourg (Lanaken--50°53N-5°39E, Vliermaalroot--50°50N-5°23E, Canne), Brabant (Louvain--50°52N-4°42E, Uccle--50°49N-4°21E, Overijse--50°46N-4°34E, Halle--50°44N-4°14E, Boistfort, Hoeilart, Bois de la Cambre, Etterbeek, Auderghem, Rouge-Cloître, Watermael, Boendael, Notre-Dame-au-Bois, Linkebeek, Rhode-Sainte-Genèse, Buysinghen, Assche), Liège (Wandre--50°40N-5°40E, Liège--50°38N-5°35E, Jupille--50°38N-5°38E, Chênée--50°37N-5°37E, Chaudfontaine--50°34N-5°40E, Huy--50°31N-5°15E, Sart--50°31N-5°57E, Spa--50°29N-5°52E, Aywaille--50°28N-5°40E, Bois d'Aywaille--50°28N-5°40E--Kin, Fléron, Ougrée, Embourg, Chèvremont, Beaufays, Tilff, Strée, Bois l'Evèque, Val Benoît, Jemeppe, Cointe, Beyne, Méry), Hainaut (Chimay--50°03N-4°19E--Forges-lez-Chimay, Harmignies, Vergnies), Namur (Namur--50°28N-4°52E, Hastière--50°14N-4°42E, Eprave, Foy-Notre-Dame, Waulsort, Marche-les-Dames), Luxembourg (Saint-Hubert--50°02N-5°23E, Bouillon--49°47N-5°05E, Laroche, Izel, Mardesous);

    Luxembourg (Echternach--49°49N-6°26E--Berdorf);

    France : Nord (Armentières--50°41N-2°52E), Calvados (Saint Aubain--49°20N-0°23W, Pont l'Evêque--49°17N-0°11E, Bures), Aisne (Vise), Marne (Chalons-sur-Marne--48°58N-4°22E), Meuse, Yvelines (Poissy--48°55N-2°05E, Saint Germain en Laye--48°52N-2°05E, Draveil, Jubisy, Bois de Boulogne, Auteuil, Longchamps, Achères, Neuilly, Vincennes, Vésinet, Croissy, Bougival, Marly), Val D'Oise (Montmorency--48°59N-2°22E), Hauts-De-Seine (Meudon--48°50N-2°15E), Paris (Paris--48°50N-2°20E), Essonne (Brétigny-sur-Orge--48°37N-2°18E), Finistère (Benodet--47°53N-4°07W), Morbihan (Grandchamp--47°46N-2°51E), Seine-Et-Marne (Melun--48°33N-2°38E--Mée), Terr. de Belfort (Delle--47°31N-6°59E), Indre-Et-Loire (Tours--47°25N-0°42E), Loir-Et-Cher (Romorantin--47°22N-1°44E), Cher (Meillant), Côte d'Or (Petigny), Allier (Saint Ennemond, Saint Evremond, Tronçais), Saône et Loire (Les Antoines, Issy l'Evêque), Charentes Maritimes (Montguyon--45°11N-0°12W), Charente (Féolle, Floirac), Dordogne, Limousin, Loire (Saint Etienne--45°28N-4°25E), Isère, Haute-Savoie (Chamonix--45°55N-6°51E, Annecy--45°54N-6°07E), Gironde (Bordeaux--44°50N-0°37W, La Ruscade), Lot-Et-Garonne (Agen--44°11N-0°40E), Tarn-Et-Garonne (Puylaroque, Montauban--44°00N-1°12E), Aveyron (Decazeville--44°33N-2°14E, Saint-George de Luzençon), Ardèche (Casteljau, Saint-Remèze, bois de Païolive), Drôme (Saint Donat sur l'Herbasse--45°07N-4°59E), Landes (Parentis--44°21N-1°07W, Dax--43°42N-1°02W), Gers (Escorneboeuf), Pyrénées-Atlantiques (Salies de Bearn--43°29N-0°58W, Saint Jean de Luz--43°21N-1°26W, Pau--43°18N-0°25W), Hautes Pyrénées (Lourdes--43°06N-0°02E, Cauterets--42°52N-0°03W, Lespoune), Ariège (Pamiers--43°06N-1°37E, Foix--42°03N1°36E), Tarn (Grésigne), Aude (Esperanza), Pyrénées Orientales (La Massane, Ria, Banyuls), Hérault (Montpellier--43°36N-3°57E, Saint Gély, Saint Guilhem, Minerve), Bouches du Rhône (Auriol--43°22N-5°38E, Marseille--43°17N-5°24E, Géménos--43°17N-5°37E, Saint Pons), Vaucluse (Mts. du Luberon--43°50N-5°20E, Oppéde), Alpes de Haute Provence (Banon--44°02N-5°37E, Riez--43°49N-6°05E), Var (Le Luc--43°24N-6°19E, La Roquebrussane--43°20N-5°59E, Mts. Sainte Baume--43°19N-5°45E, Forêt de Sainte-Baume--43°19N-5°45E, Pignans--43°18N-6°14E, Signes--43°17N-5°50E, Montrieux le Vieux--43°15N-5°58E, Collobrières--43°15N-6°19E, Belgentier--43°14N-6°01E, Toulon--43°07N-5°55E, Hyères--43°07N-6°04E, Correns, forêt de Morrières, Evenos, Aiguines, Mt. Touar, Valbelle), Alpes Maritimes (Saint Cézaire, Gilette), Corse--42°N-9°E, ? (Guyon, Saint Martin Vesubie);

    Spain : Cantabria : Potes, 43°09 N - 4°37 W

    Spain : Guipúzcoa : Tolosa, 43°09 N - 2°05 W

    Spain : Cataluña

    Spain : Huesca : Torla, 42°38 N - 0°06 W

    Spain : Huesca : Benasque, 42°36 N - 0°31 W

    Spain : Huesca : Valle de Gistain, 42°36 N - 0°20 E

    Spain : Huesca : Plan, 42°35 N - 0°20 E

    Spain : Huesca : Jaca, 42°34 N - 0°33 W

    Spain : Huesca : Loarre, 42°19 N - 0°37 W

    Spain : Huesca : La Peña

    Spain : Teruel : Albarracín, 40°30 N - 1°34 W

    Germany : Hamburg (Hamburg--53°32N-10°00E), Nordrhein-Westfalen (Dortmund--51°31N-7°27E), Hessen (Wetzlar--50°33N-8°31E, Kassel--51°19N-9°28E), Rheinland (Koblenz--50°22N-7°36E), Baden-Wurtemberg (Pforzheim--48°55N-8°42E), Rostock, Schwerin, Neubrandenburg, Magdeburg, Potsdam (Velten--52°41N-13°11E), Berlin, Frankfurt, Halle, Cottbus, Erfurt, Leipzig, Dresden, Suhl, Gera, Chemnitz = Karl-Marx-Stadt, ? (Kauden, Bentheumer Woud);

    Poland;

    Czech Rep. : Vinnéjezero, Pálava Biosphere Reserve;

    Slovakia;

    Switzerland : Basel : Kleinhuningen, 47°35 N - 7°35 E, 252 m, 1955 (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Basel : Riehen, 47°35 N - 7°39 E : Schaferstrasse, 281 m, 1988 (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Basel : Bachgraben, 275 m (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Basel : Baumlihof, 259 m (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Basel : Lange Erlen, 253 m, 1965 (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Basel : Schwarz-Park, 261 m, 1992 (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Basel : Bettingen, 47°34 N - 7°40 E : Chrischona, 470 m (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Basel : Basel, 47°33 N - 7°35 E (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Augst, 47°32 N - 7°43 E : Schlossli, 274 m, 1989 (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Augst, 47°32 N - 7°43 E : Schonbuhl, 289 m, 1965 (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Munchenstein, 47°31 N - 7°37 E : Hofmatt, 273 m, 1984 (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Munchenstein, 47°31 N - 7°37 E : Ob. Wannen, 332 m, 1990 (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Muttenz, 47°31 N - 7°39 E : Schanzli, 260 m, 1991 (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Muttenz, 47°31 N - 7°39 E : Rutihard, 370 m, 1990 (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Giebenacht, 47°31 - 7°48, 318 m, 1992 (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Therwil, 47°30 N - 7°33 E : Froloo, 391 m, 1942 (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Reinach, 47°30 N - 7°35 E : Leiwald, 310 m (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Arlesheim, 47°30 N - 7°37 E : Spitalholz, 348 m, 1990 (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Arlesheim, 47°30 N - 7°37 E : Schwimmbad, 300 m (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Arlesheim, 47°30 N - 7°37 E : Reichenstein, 460 m (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Lausen, 47°29 N - 7°45 E, 344 m, 1984 (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Blauen, 47°27 N - 7°31 E : Weekendhauser, 570 m (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Bubendorf, 47°27 N - 7°44 E : Wildenstein, 475 m (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Zwingen, 47°26 N - 7°32 E : Bahnhof, 339 m, 1995 (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Himmelried, 47°26 N - 7°35 E: Steffen, 574 m, 1995 (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Windentalhohe, 420 m, 1989 (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Grammont, 539 m, 1989 (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Schleifenberg, 534 m, 1990 (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Bern : Uetental, 380 m (citated by Sprecher-Uebersax & Durrer, 1998).

    Switzerland : Jura : Lajoux, 47°17 N - 7°08 E

    Switzerland : Vaud : Genève, 46°12N-6°09E

    Switzerland : Bagatz

    Osterreich : Ost-Tirol : Thuringen, 47°12 N - 9°46 E

    Osterreich : Donau : Neuburg Bergheim

    Osterreich : Wien, 48°12 N-16°21 E

    Hungary : Mt. Bukk, 48°04N-20°29E

    Hungary : Gödölló, 47°37N-18°54E

    Hungary : Veszprém, 47°05N-17°57E

    Hungary : Lake Balaton, 46°50N-17°45E : Tihany

    Hungary : Mecsek Mts., 46°10N-17°56E

    Hungary : Zebegewy

    Hungary : Pomaz

    Hungary : Borosjenö

    Hungary : Torokbalint

    Romania

    Italy : Piemonte (Arona--45°45N-8°32E, Biella--45°34N-8°01E), Lombardia (Varese--45°49N-8°51E--Montvalle, Como--45°48N-9°05E), Venezia Giulia (Udine--46°05N-13°12E, Udine--46°05N-13°12E--Toppo, Vittorio--45°58N-12°17E, Pordenone--45°57N-12°38E, Treviso--45°39N-12°14E, Trieste--43°40N-13°46E {San Sergio-Petrigna, Valle Ostra}), Liguria (Genova--44°22N-8°55E), Toscana (Firenze--43°47N-11°16E = Florence, Siena--43°19N-11°19E, Cortona--43°15N-12°00E, Badia Prataglia), Emilia Romagna (La Lama), Sardegna--40°10N-9°E, Sicilia--37°35N-14°15E, ? (Zattaglia);

    Slovenija: Mackovci--46°47N-16°10E, Maribor--46°34N-15°37E, Celje--46°15N-15°14E, Celje--46°15N-15°14E--Ljubno ob Savinja River, Ljubljana--46°04N-14°29E, Litija--46°03N-14°52E--Hotic, Grosuplje--45°57N-14°40E--Tabor, Ajdovscina--45°53N-13°53E, Cerknica--45°48N-14°20E, Postojna--45°47N-14°11E--Javorniki, Sezana--45°43N-13°50E, Kocevje--45°39N-14°49E, Kozina--45°34N-13°56E, Bohinj--Ribcev laz, Bohinj--Vogar, ob Suhi-Bohinj, Bohinjska Bistrica, Crna prst, Gameljne, Gorica, Kamnik, Komenda, Kostanjevica, Kuzma, Lubnik, Orehovci--Gornja Radgona, Pekel--Zalec, Podgorje--Slavnik, Radenci, Rakek, Rakov Skocjan, Senozece, Sorica, Sorsko polje, Tacen, Tatre, Trzin, Vaneca, Vipava, Vodice, Zg. Tuhinj;

    Bosnia & Hercegovina : Hercegovina (Konjice--43°39N-17°56E, Mostar--43°20N-17°48E);

    Yugoslavia : ? (Karlovak);

    Bulgaria;

    Greece : Karpenision--38°55N-21°46E;

    Turkey;

    Russia : to Ural and Caucasus (Terek River, Khosta);

    Iran.

    Note : this species is protected by Bern Convention appendix 3; Habitat Direction appendix 2; Regional Law of Toscana, Italia, appendix A and B (Ballerio, 2003).

    Xylophagous :

    Betulaceae : Alnus, Carpinus.

    Fagaceae : Quercus, Fagus.

    Hippocastanaceae : Aesculus.

    Juglandaceae : Juglans.

    Malaceae : Pyrus, Malus.

    Moraceae : Morus.

    Oleaceae : Fraxinus.

    Pinaceae : Picea, Pinus.

    Rosaceae : Cerasus, Prunus.

    Salicaceae : Salix, Populus.

    Tiliaceae : Tilia.

    Ulmaceae : Ulmus.

    Saproxylophagous :

    Aceraceae : Acer.

    Fagaceae : Quercus.

    Can breed in compost (artificial soil).

    Natural enemies :

    Birds : Corvidae : Pica.

    ? : Caprimulgus europaeus, Strix aluco.

    Acarina : Uropodidae : Polyaspis patavinus.

    ? : Isacis cuspidat, Isacis lucani, Megaselia rufipes, Monocercomonas melolonthae.

    Bibliografía :

    BAGUENA L. (1967) Scarabaeoidea de la fauna íbero-balear y pirenaica. C.S.F.C. Madrid, 1955 (publicado en 1967): 1-567.

    BALLERIO A. (2003) EntomoLex : la conservazione degli insetti e la legge. Memorie Soc. entomol. Italiana, 82(1):17-86.

    COFAIS M. (1972) Un cas tératologique chez Lucanus cervus. L'Entomologiste, 28:32-33.

    CLARK J.T. (1977) Aspects of variation in the sag beetle. Systematic Entomology, 2:9-16.

    FREUDE H. (1968) Uber Mandibel-Anomalitaten bei Lucanus cervus. Nachrichtenblatt der Bayerischen Entomologen, 17:59-64.

    HORION A. (1958) Faunistik der mitteleuropaischen Kafer 6: Lamellicornia. 343 pp.

    KLAUSNITZER B. (1995) Die Hirschkafer. Neue Brehm-Bucherei, 551:109 pp.

    LOPEZ-COLON J.I., GONZALEZ PEÑA C.F. & BELTRAN VALEN J.R. (1996) Familia : Lucanidae. Catalogus de la Entomofauna Aragonesa, No. 12:15-20.

    MAL N. (1972) Sur le polymorphisme antennaire d'une population provençale de Lucanus cervus. L'Entomologiste, 38:161-166.

    OCHSE M. (1993) Angaben zum Hirschkafer, Lucanus cervus L., bei Kassel. Hessische faunistische Briefe, 13:47-52.

    REYES-CASTILLO P. (2002) 27. Lucanidae. Pp. 485-492. En LLORENTE BOUSQUETS J. & MORRONE J.J. Biodiversidad, Taxonomía y Biogeografía de Artrópodos de México : Hacia una síntesis de su conocimiento. Universidad Nacional Autónoma de México. Vol. III:692 pp.

    ROER H. (1980) Zur Verbreitung und Bestandesdichte des Hirschkafers im Rheinland. Acta Musei Rigenaehradecensis, Serie A, Supplement: 248-251.

    SPRECHER-UEBERSAX E. & DURRER H. (1998) Uber das Vorkommen des Hirschkafers (Lucanus cervus L.) in der Region Basel (Coleoptera). Mitt. Ent. Ges. Basel, 48(4):142-166.

    SPRECHER-UEBERSAX E. & DURRER H. (1998) Untersuchungen zum Stridulationsverhalten der Hirschkafer-Larven (Lucanus cervus L.) (Coleoptera : Lucanidae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 71:471-479.

    SPRECHER-UEBERSAX E. & DURRER H. (2001) Beobachtungen zur Nahrungswahl des Hirschkafers (Lucanus cervus L.) (Coleoptera). Mitteilungen der Entomologischen Gesellschaft Basel, 51(1):2-11.

    SPRECHER-UEBERSAX E. & DURRER H. (2001) Verhaltenstudien beim Hirschkafers mittels Telemetrie und Videoaufzeichnungen (Coleoptera, Lucanus cervus L.). Mitteilungen der Naturforschenden Gesellschaften beider Basel, 5:161-182.

    ZEMP M., HARTMANN K., KNECHT D. & SPRECHER-UEBERSAX E. (1990) Arlesheimer Wald Spitalholz 1990. Unveroffentliches Manuskript zuhanden der Kober-Schwabe-Stiftung Arlesheim, 59 pp.

     

    TEXT : MAES JEAN-MICHEL 

     

    PICTURES : GALANT ALAIN

     

  • cervus ssp. fabiani Mulsant & Godart, 1855

    4 photos

     

    Lucanus (Lucanus) cervus fabiani

     

     

    Lucanus (Lucanus) cervus ssp. fabiani MULSANT & GODART, 1855 [Platycerus].

    Hexaphyllus pontbrianti MULSANT 1839 [Lucanus, Platycerus] [France : Lyon : Bois de Rochecardon].

    Lucanus pentaphyllus REICHE 1853 [Platycerus] [Gallia meridionalia : circa Telonem et Portum Veneris].

    Lucanus heptaphyllus ABEILLE DE PERRIN 1867.

    Lucanus reichei MOTSCHULSKY 1870 [France : Toulouse].

    Lucanus capreolus hexaphyllus (PLANET) DIDIER & SEGUY 1953 [Italie] (nomen nudum).

    SE. France : Rhône (Lyon--45°45N-4°51E--bois de Rochecardon), Ardèche, Drôme (Nyons--44°21N-5°08E), Landes (Dax--43°42N-1°02W), Pyrénées Orientales (Prades--42°37N-2°25E, Port-Vendres--42°29N-3°07E, Ria, Vernet-les-Bains, Banyuls-sur-Mer), Hérault (Montpellier--43°36N-3°57E), Var (La Roquebrussane--43°20N-5°59E, Montrieux le Vieux--43°15N-5°58E, Toulon--43°07N-5°55E, Hyères--43°07N-6°04E), Alpes Maritimes, Corse (St. Hyacinthe, Ponte Vecchiu near Ota).

    Note : For Lacroix (1968) fabiani is only a morfa of cervus because cervus and fabiani are living in the same region. 

     

    TEXT : MAES JEAN-MICHEL

     

    PICTURES : GALANT ALAIN

     

  • cervus ssp. judaicus Planet, 1902

    3 photos

     

    Lucanus (Lucanus) cervus judaicus

     

     

    Lucanus (Lucanus) cervus ssp. judaicus PLANET 1902.

    Turkey : Aïn-Tab* at 90 km NE Alep.

     

    TEXT : MAES JEAN-MICHEL 

     

    PICTURES : GALANT ALAIN

     

  • cheni Huang, He & Shi, 2011

    5 photos

    Lucanus cheni Huang, He & Shi Z., 2011.

     

    China - Xizang (Tibet)

     

    HUANG, HE & SHI, 2011. Description of two new species of Lucanus Scopoli from southeastern Tibet, China; Zootaxa 2987:56-64.

     

     

    TEXT : GALANT ALAIN 

     

    PICTURES : GALANT ALAIN

  • cyclommatoides Didier, 1928

    4 photos

     

    Lucanus (Lucanus) cyclommatoides

     

     

    Lucanus (Lucanus) cyclommatoides DIDIER, 1928.

    Lucanus cyclommatoides DIDIER, 1928:51

    Lucanus cyclommatoides DIDIER; Didier & Séguy, 1953:81

    Lucanus cyclommatoides DIDIER; Benesh, 1960:140

    Lucanus cyclommatoides DIDIER; Maes, 1992:17.

    Lucanus cyclommatoides DIDIER; Araya, 2001:12, figs.16-20.

    •Vietnam : Tonkin : Chapa, 13-VII, col. Jeanvoine (male TYPE of Lucanus cyclommatoides DIDIER, 1928 en Museum National d'Histoire Naturelle, Paris).
    •Vietnam : Tonkin : Chapa, 26-IX-1918, col. Jeanvoine, ex. coll. Boileau (female COTYPE of Lucanus cyclommatoides DIDIER, 1928 en Museum National d'Histoire Naturelle, Paris).

     

    Bibliography :

    ARAYA K. (2001) Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae) from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.

     

    TEXT : MAES JEAN-MICHEL

     

    PICTURES : GALANT ALAIN

     

  • datunensis Hashimoto, 1984

    4 photos

     

    Lucanus (Lucanus) datunensis

     

     

    Lucanus (Lucanus) datunensis HASHIMOTO, 1984.

    Lucanus datunensis HASHIMOTO, 1984:18 [Taiwan : Mt. Datun].

    Lucanus ritae LACROIX, 1984:10 [Mt. Ali-shan].

    Lucanus ritae LACROIX; Lacroix, 1984:5.

    Lucanus datunensis HASHIMOTO; Sakaino, 1985:15-18

    Lucanus datunensis HASHIMOTO; Nagai, 1985.

    Lucanus datunensis HASHIMOTO; Kawano, 1988:26

    Lucanus datunensis HASHIMOTO; Hashimoto, 1990:23-26, pl.4

    Lucanus datunensis HASHIMOTO; Wang, 1990.

    Lucanus datunensis HASHIMOTO; Tanikado, 1992.

    Lucanus datunensis HASHIMOTO; Chang, 1993.

    Lucanus datunensis HASHIMOTO; Sakaino & Yu, 1994:26-27

    Lucanus datunensis HASHIMOTO; Sakaino, Yu & Chu, 1994:122

    Lucanus datunensis HASHIMOTO; Mizunuma & Nagai, 1994:217, pl.16.

    Lucanus datunensis HASHIMOTO; Wang, 1994.

    Lucanus datunensis HASHIMOTO; Araya, 2000:1-4.

    Lucanus ritae LACROIX; Araya, 2001:19-20, fig.71.

    Lucanus datunensis HASHIMOTO; Araya, 2001:19.

    •Taiwan : Taipei Distr. : Mt. Tatun Shan = Mt. Datun (male HOLOTYPUS of Lucanus datunensis HASHIMOTO, 1984).
    •Taiwan : Central Taiwan : Mt. Ali Shan (male HOLOTYPUS of Lucanus ritae LACROIX, 1984) (2 males PARATYPUS of Lucanus ritae LACROIX, 1984 in Museum d'Histoire Naturelle de Geneve). Bibliography.
    ARAYA K. (2000) Notes on Lucanus ritae Lacroix, 1984, a junior synonym of Lucanus datunensis Hashimoto, 1984. Kogane, Tokyo, 1:1-4.
    ARAYA K. (2001) Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae) from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
    CHANG Y. (1993) Stag beetles of Taiwan. Newton Press, Taipei, 111 pp. [in Chinese].
    HASHIMOTO S. (1984) A new species of the genus Lucanus of the family Lucanidae inhabiting Taiwan. Iwase, 2:23-26.
    LACROIX J.P. (1984) Descriptions de Coleoptera Lucanidae nouveaux ou peu connus. Bull. Sciences Nat, 40(1983):5-18, 7 pls.
    MIZUNUMA T. & NAGAI S. (1994) The Lucanidae of the world. Mushi-sha's Iconographic Series of Insects. 1:337 pp. Mushi-sha, Tokyo.
    NAGAI S. (1985) Coleopterorum Catalogus, World Lucanidae. Mokuyo-sha, Tokyo. 224 pp. [In Japanese].
    SAKAINO H. (1985) Miscellaneous notes on a Formosan Lucanidae, Lucanus datunensis Hasimoto. Gekkan-Mushi, 172:15-18.
    SAKAINO H. & YU C.K. (1994) The female of Lucanus datunensis Hashimoto (Coleoptera, Lucanidae). Gekkan-Mushi, 285:26-27.
    TANIKADO M. (1992) Current topics of stag beetles in Taiwan. Gekkan-Mushi, 256:22-25.
    WANG H.Y. (1990) Illustrations of stag beetles in Taiwan. Taiwan Museum, Taipei, 189 pp. (In Chinese).
    WANG H.Y. (1994) Illustrations of stag beetles in Taiwan. Taiwan Museum, Taipei, 207 pp. (In Chinese).

     

     

    TEXT : MAES JEAN-MICHEL 

     

    PICTURES : GALANT ALAIN

     

  • delavayi Fairmaire, 1887

    8 photos

     

    Lucanus (Lucanus) delavayi

     

     

    Lucanus (Lucanus) delavayi FAIRMAIRE, 1887.

    Lucanus delavayi FAIRMAIRE, 1887:27 [China : Yunnan].

    Lucanus delavayi FAIRMAIRE; Fairmaire, 1887:98

    Lucanus delavayi FAIRMAIRE; Planet, 1900:228

    Lucanus delavayi FAIRMAIRE; Planet, 1902:68

    Lucanus delavayi FAIRMAIRE; Heyne & Taschenberg, 1908:50

    Lucanus delavayi FAIRMAIRE; Didier, 1949:pl.10

    Lucanus delavayi FAIRMAIRE; Benesh, 1960:140

    Lucanus delavayi FAIRMAIRE; Kurosawa, 1978:143

    Lucanus delavayi FAIRMAIRE; Bomans, 1989:9

    Lucanus delavayi FAIRMAIRE; Maes, 1992:17

    Lucanus delavayi FAIRMAIRE; Mizunuma & Nagai, 1994:216, pl.16.

    Lucanus delavayi FAIRMAIRE; Araya, 2001:13, figs.25-26.

    China : Yunnan, col. R.P. Delavay (male PARATYPE of Lucanus delavayi FAIRMAIRE, 1887, # 1666 in Museum d'Histoire Naturelle, Geneva).
    China : Yunnan : Ta-Pin-Tze
    China : Yunnan : Yunnan-Sen
    China : Yunnan : San-Nen-Kai.

     

    Bibliography :
    ARAYA K. (2001) Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae) from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22. 

     

    TEXT : MAES JEAN-MICHEL

     

    PICTURES : GALANT ALAIN

     

  • dohertyi Boileau, 1911

    5 photos

     

    Lucanus (Lucanus) dohertyi

     

     

    Lucanus (Lucanus) dohertyi BOILEAU, 1911.

    Lucanus laminifer var. minor WATERHOUSE 1890.

    Waterhouse, 1890:33; Boileau, 1911:435, pl.34; Arrow, 1950:56; Didier & Séguy, 1952:pl.15; Benesh, 1960:140; Lacroix & Bomans, 1973:138, 142; Lacroix, 1982:13; Téllez, 1990:39; Maes, 1992:17; Mizunuma & Nagai, 1994:214, 309, pl.11.

    India : Assam, Meghalaya (Khasi Hills), Nagaland (Naga Hills*).

     

    TEXT : MAES JEAN-MICHEL 

     

    PICTURES : GALANT ALAIN

     

  • dongi Okuda, 2009

    4 photos

     

    Lucanus (Lucanus) dongi

     

    Lucanus dongi , MAEDA T. 2009.

    Ø       Central Vietnam

                                                                                                                                                

    Bibliography :

     

    MAEDA T. ( 2009 ) Three new species of the genera Lucanus, Rhaetulus and Dorcus ( Coleoptera, Lucanidae ) from central Vietnam. Gekkan-Mushi, 2009; N°457, Mar. 2009; p 35-70. 

     

     

    TEXT : GALANT ALAIN

     

    PICTURES : GALANT ALAIN

  • elaphus Fabricius, 1775

    5 photos

     

    Lucanus (Lucanus) elaphus

     

     

    Lucanus (Lucanus) elaphus FABRICIUS, 1775.

    Lucanus cervus var. DEGEER 1774:33.

    Lucanus elaphus FABRICIUS, 1775:2 [Virginia].

    Lucanus elaphus FABRICIUS; Fabricius, 1781:2.

    Lucanus elaphus FABRICIUS; Fabricius, 1887:1.

    Lucanus elaphus FABRICIUS; Olivier, 1789:12-13, pl.3.

    Lucanus elaphus FABRICIUS; Linnaeus (ed. Gmelin), 1790:1589.

    Lucanus elaphus FABRICIUS; Fabricius, 1792:237.

    Lucanus elaphus FABRICIUS; Olivier (ed. Illiger), 1800:60-61.

    Lucanus elaphus FABRICIUS; Fabricius, 1801:249.

    Lucanus elaphus FABRICIUS; Olivier (ed. Sturm), 1802:2, pl.2.

    Lucanus elaphus FABRICIUS; Thunberg, 1806:191.

    Lucanus elaphus FABRICIUS; Schonherr, 1817:322.

    Lucanus elaphus FABRICIUS; Dejean, 1837:193.

    Lucanus elaphus FABRICIUS; Burmeister, 1847:354.

    Lucanus elaphus FABRICIUS; Fuchs, 1882:49.

    Lucanus elaphus FABRICIUS; Albers, 1890:88.

    Lucanus elaphus FABRICIUS; Planet, 1902:92, pl.16.

    Lucanus elaphus FABRICIUS; Ulke, 1902.

    Lucanus elaphus FABRICIUS; Wickham, 1903:205-206.

    Lucanus elaphus FABRICIUS; Heyne & Taschenberg, 1908:50, pl.6.

    Lucanus elaphus FABRICIUS; Blatchley, 1910:905.

    Lucanus elaphus ab. carlengi ANGELL 1916:70 [Kentucky : Louisville].

    Lucanus elaphus FABRICIUS; Brimley, 1938.

    Lucanus elaphus FABRICIUS; Loding, 1945.

    Lucanus elaphus FABRICIUS; Didier, 1949:pl.11.

    Lucanus elaphus FABRICIUS; Benesh, 1960:140-141.

    Lucanus elaphus FABRICIUS; Arnett, 1965.

    Lucanus elaphus FABRICIUS; Ritcher, 1966.

    Lucanus elaphus FABRICIUS; Kirk, 1969.

    Lucanus elaphus FABRICIUS; Kirk, 1970.

    Lucanus elaphus FABRICIUS; Arnold & Drew, 1987.

    Lucanus elaphus FABRICIUS; McNamara, 1991.

    Lucanus elaphus FABRICIUS; Ratcliffe, 1991:267.

    Lucanus elaphus FABRICIUS; Maes, 1992:17.

    Lucanus elaphus FABRICIUS; Anonymous, 1994.

    Lucanus elaphus FABRICIUS; Mizunuma & Nagai, 1994:212, pl. 7.

    Lucanus elaphus FABRICIUS; Staines, 1994:17-18.

    Lucanus elaphus FABRICIUS; Poole & Gentili, 1996:297.

    Lucanus elaphus FABRICIUS; Staines, 2000:26-30.

    Lucanus elaphus FABRICIUS; Staines, 2001:397-404.

    Canada :

    Ontario (citated by Benesh, 1960).

    USA :

    Minnesota (National Museum of Natural History, Washington).

    Michigan (Ohio State University).

    Nebraska : Richardson Co. : Stella, 6-VI-1999 (University of Nebraska).

    Iowa : Lee Co. : Keokuk (Museum of Comparative Zoology, Cambridge).

    Kansas (Field Museum of Natural History, Museum of Comparative Zoology, Cambridge, National Museum of Natural History, Washington).

    Kansas : Douglas Co. (University of Kansas).

    Kansas : Labette Co. : Oswego (National Museum of Natural History, Washington).

    Kansas : Leavenworth Co. : Leavenworth, 23-VII-1943 (California Academy of Science).

    Kansas : Leavenworth Co. : Leavenworth, 30-VII-1947 (University of Kansas).

    Kansas : Wilson Co. : New Albany (Natural History Museum, London).

    Kansas : Wyandotte Co. (Field Museum of Natural History).

    Missouri (Field Museum of Natural History).

    Missouri : Boone Co. : Columbia, VI-1903 (Field Museum of Natural History).

    Missouri : Boone Co. : Columbia, 21-VI-1971 (University of Missouri).

    Missouri : Boone Co. : Columbia, 14-VII-1959 (University of Missouri).

    Missouri : Boone Co. : Columbia, 2-VIII-1968 (University of Missouri).

    Missouri : Boone Co. : Columbia, 4-IX-1965 (University of Missouri).

    Missouri : Butler Co. : 20-VI-1897 (National Museum of Natural History, Washington).

    Missouri : Poplar Bluff, VII-1952 (University of Missouri).

    Missouri : Girardeau Co. : Cape Girardeau, 25-VII-1956 (National Museum of Natural History, Washington).

    Missouri : Jasper Co. (IMRM).

    Missouri : Jefferson Co. : Barnhart, 30-VII-1960 (University of Missouri).

    Missouri : Jefferson Co. : Barnhart, 3-VIII-1935 (University of Missouri).

    Missouri : New Madrid Co. : Morehouse, 6-V-1915 (Museum of Comparative Zoology, Cambridge).

    Missouri : Pike Co. : VII-1934 (University of Missouri).

    Missouri : Louisiana, 6-VI-1923 (University of Missouri).

    Missouri : Platte Co. : 29-VI-1992 (University of Missouri).

    Missouri : Pulaski Co. : 17-VII-1956 (Florida State Collection of Arthropods).

    Missouri : St. Charles Co. : O'Fallen, 4-VI-1904 (University of Missouri).

    Missouri : St. Louis Co. : Creve Coeur, 1-VII-1908 (University of Missouri).

    Missouri : Meramec Hinds, 3-VI-1923 (National Museum of Natural History, Washington).

    Missouri : St. Louis, 20-VII-1897 (National Museum of Natural History, Washington).

    Missouri : St. Louis, 20-VI-1904 (Field Museum of Natural History).

    Missouri : St. Louis, 23-VI-1907 (University of Michigan).

    Missouri : St. Louis, 28-VI-1972 (University of Missouri).

    Missouri : St. Louis (California Academy of Science, Carnegie Museum of Natural History).

    Missouri : Stoddard Co. : Popular Bluff Forestry Camp, 23-VI-1954, taken at light (Florida State Collection of Arthropods).

    Missouri : Stone Co. : 20-VIII-1915 (University of Missouri).

    Missouri : Taney Co. : Forsyth, 14-VII-1929 (Field Museum of Natural History).

    Missouri : Texas Co. : Mountain Grove, 1916 (National Museum of Natural History, Washington).

    Missouri : Washington Co. : Irondale, 29-VII-1961 (University of Missouri).

    Missouri : Wayne Co. : Williamsville, 20-III-1961 (University of Missouri).

    Missouri : Wright Co. : Cedar Gap, 5-VI-1911 (Field Museum of Natural History).

    Illinois (University of Minnesota).

    Illinois : Southern Illinois (National Museum of Natural History, Washington).

    Illinois : Atlas, VI-1931 (University of Missouri).

    Illinois : Adams Co. : Quincy (Illinois Natural History Survey).

    Illinois : Alexander Co. : Cairo (National Museum of Natural History, Washington).

    Illinois : Villa Ridge, IX-1892 (Illinois Natural History Survey).

    Illinois : Calhoun Co. : Batchtown, 8-VI-1899 (Illinois Natural History Survey).

    Illinois : Clay Co. : Flora (National Museum of Natural History, Washington).

    Illinois : Edwards Co. : 15-VI-1944 (Illinois Natural History Survey)

    Illinois : Hamilton Co. : Dahlgren, 4-VI-1885 (Illinois Natural History Survey).

    Illinois : Knox Co. : Galesburg (Museum of Comparative Zoology, Cambridge)

    Illinois : Platt Co. : Allerton Estate, 26-VI-1959 (Illinois Natural History Survey).

    Illinois : Randolph Co. : 24-V-1896 (University of Missouri).

    Illinois : Richland Co. : 3-VI-1926 (Illinois Natural History Survey).

    Illinois : Wabash Valley, V-1878 (Museum of Comparative Zoology, Cambridge)

    Illinois : St. Clair Co., 21-VI-1902 (University of Missouri)

    Illinois : St. Clair Co., 1-VIII-1902 (University of Missouri)

    Illinois : Manssa, 30-V-1921 (Illinois Natural History Survey).

    Illinois : Union Co. : Anna, 17-VII-1883 (Illinois Natural History Survey)

    Illinois : Pine Hills Recreation Area, 10-VII-1968, blacklight trap (Florida State Collection of Arthropods).

    Illinois : Washington Co. : Dubois, 1909 (Illinois Natural History Survey).

    Indiana (National Museum of Natural History, Washington).

    Indiana : Clay Co. : Saline City, 6-VI-1889 (National Museum of Natural History, Washington).

    Indiana : Crawford Co. (citated by Blatchley, 1910).

    Indiana : Daviess Co. : Bicknell, 4-VI-1937 (University of Arkansas).

    Indiana : Daviess Co. : Bicknell,  6-VI-1938 (University of Arkansas).

    Indiana : Dubois Co. : 13-VIII-1970 (Texas A & M University).

    Indiana : Elkhart Co. : Elkhart (California Academy of Science).

    Indiana : Knox Co., 27-V-1944 (University of Arkansas).

    Indiana : Knox Co., 10-VI-1943 (University of Arkansas).

    Indiana : Monroe Co. : Bloomington, VI-1928 (California Academy of Science)

    Indiana : Monroe Co. : Bloomington, 13-VI-1931 (California Academy of Science)

    Indiana : Orange Co. : Orleans, 3-VII-1935 (University of Arkansas).

    Indiana : Vigo Co. (citated by Blatchley, 1910).

    Indiana : Wabash Co. : Mt. Vernon, 23-V-1964 (Florida State Collection of Arthropods).

    Indiana : Corydon (in coll. Jay Timberlake, USA).

    Ohio (National Museum of Natural History, Washington).

    Ohio : Bowlin, VII-1937 (Ohio State University).

    Ohio : Erie Co. : Sandusky (Cedar Point), 23-VII-1904 (California Academy of Science).

    Ohio : Hamilton Co. : Cincinnati (National Museum of Natural History, Washington).

    Kentucky (Illinois Natural History Survey).

    Kentucky : Cumberland Gap (Museum of Comparative Zoology, Cambridge).

    Kentucky : Oragus (Museum of Comparative Zoology, Cambridge).

    Kentucky : Bell Co. : Balkan, VI-1915 (National Museum of Natural History, Washington).

    Kentucky : Henderson Co. : Henderson, 25-V-1921 (Field Museum of Natural History).

    Kentucky : Jefferson Co. : Louisville, 38°15N-85°45W, 9-VII-1938 (University of Kentucky)

    Kentucky : Letcher Co. : Whitesburg, VIII-1915 (National Museum of Natural History, Washington).

    Kentucky : Meade Co. : Otter Creek Park, 19-VI-1970 (National Museum of Natural History, Washington).

    Kentucky : Todd Co. : Elkton, 17 June 1961 (University of Kentucky).

    Kentucky : Warren Co. : Bowling Green, VII-1892 (University of Kentucky).

    Pennsylvania (Field Museum of Natural History).

    Pennsylvania : Delaware Co. : Lansdowne, 10-VI-1935 (California Academy of Science).

    Pennsylvania : Greene Co. : Mapletown, 4-I-1961 (National Museum of Natural History, Washington).

    Pennsylvania : Montgomery Co. (Museum of Comparative Zoology, Cambridge).

    West Virginia : Logan Co. : Fort Branch, 27-V-1911 (National Museum of Natural History, Washington).

    West Virginia : Marion Co. : Fairmont, 1926 (Illinois Natural History Survey).

    West Virginia : Mingo Co. : Laurel Creek Public Hunting and Fishing Area, 14-VI-1979 (West Virginia University).

    Virginia (TYPUS de Lucanus elaphus FABRICIUS)

    Virginia : Buckingham Co. : 27-V-1911 (National Museum of Natural History, Washington).

    Virginia : Lunenburg Co. : Victoria (National Museum of Natural History, Washington).

    Virginia : Montgomery Co. : Blacksburg, 5-V-1956 (Virginia Polytechnic Institute).

    Virginia : Nelson Co., 9-VII-1914 (National Museum of Natural History, Washington).

    Virginia : Nelson Co., 5-VIII-1927 (National Museum of Natural History, Washington).

    Virginia : Pittsylvania Co. : Chatham, 26-VII-1939 (Virginia Polytechnic Institute).

    Virginia : Danville (Field Museum of Natural History).

    Virginia : Prince Edward Co. : Farmville, 24-V-1923 (University of Georgia).

    Virginia : Prince Edward Co. : Farmville, 24-VI-1924 (University of Georgia).

    Virginia : Prince George Co. : Hopewell, 23-VI-1930 (National Museum of Natural History, Washington).

    Virginia : Petersburg Fort Lee, 23-VI-1962 (National Museum of Natural History, Washington)

    Virginia : 10 miles south of Petersburg, 4-VI-1970 (National Museum of Natural History, Washington).

    Virginia : Richmond Co. : Warsaw, 16-VI-1938 (National Museum of Natural History, Washington).

    Virginia : Sussex Co. : Jarratt, 18-VI (Cornell University).

    District of Columbia (citated by Uike, 1902)

    Maryland : Baltimore (Natural History Museum, London).

    Maryland : St. Mary's Co. : Great Mills, 18-VIII-1947, taken at light (Florida State Collection of Arthropods)

    Maryland : St. Mary Co. : Lexington Park, 4-VII-1992, taken at light (Florida State Collection of Arthropods)

    Maryland : Patuxent River Naval Air Station, 7-VIII-1975 (Philip J. KEAN, USA).

    Maryland : Worcester Co. : Pocomoke City, taken in blacklight trap (Florida State Collection of Arthropods).

    Maryland : Worcester Co. : Pocomoke City, 1896 (University of Maryland).

    Delaware : Sussex Co. : Milton, 6-VI-1940 (University of Delaware).

    Oklahoma : Choctow Co. : Boswell, 3-VI-1925 (California Academy of Science).

    Oklahoma : Craig Co. : Blue Jacket, 30-VI-1931 (Field Museum of Natural History).

    Oklahoma : Creek Co. : Dumright, 13-VI-1925 (California Academy of Science).

    Oklahoma : Latimer Co., VII-1982 (Florida State Collection of Arthropods).

    Oklahoma : Latimer Co., VII-1987 (Florida State Collection of Arthropods).

    Oklahoma : Red Oak, VII-1981 (Florida State Collection of Arthropods).

    Oklahoma : 5 miles west of Red Oak, 16-VII-1977 (Florida State Collection of Arthropods).

    Oklahoma : 5 miles west of Red Oak, VII-1979 (Florida State Collection of Arthropods).

    Oklahoma : Mays Co., 12-VII-1980 (Texas A & M University).

    Oklahoma : McCurtain Co., 8-VI-1934 (Field Museum of Natural History).

    Oklahoma : McCurtain Co., 11-VII-1932 (Field Museum of Natural History).

    Oklahoma : 6.5 miles northeast of Idabel, 28/29-VIII-1937 (University of Michigan).

    Oklahoma : Mclntosh Co. (citated in Arnold and Drew, 1987).

    Oklahoma : Okmulgee Co. : Haskell, VII-1905 (National Museum of Natural History, Washington).

    Oklahoma : Osage Co. : Bartlesville (National Museum of Natural History, Washington).

    Oklahoma : Ottawa Co. : Commerce, VIII-1934 (University of Kansas)

    Oklahoma : Wyandotte, 15-VI-1931 (University of Kentucky).

    Oklahoma : Payne Co. : Still-water, 18-VII-1928 (California Academy of Science).

    Arkansas : Arkansas Co. : 8-VI-1970, blacklight (University of Arkansas).

    Arkansas : Ashley Co. : Hamburg (University of Arkansas).

    Arkansas : Benton Co. : Siloam Springs, 18-VI-1907 (National Museum of Natural History, Washington).

    Arkansas : Columbia Co. : Magnolia, 1931 (University of Kansas).

    Arkansas : Devil's Den State Park, 20-VI-1975 (University of Arkansas).

    Arkansas : Fayetteville (University of Arkansas).

    Arkansas : Garland Co. : Hot Springs (National Museum of Natural History, Washington).

    Arkansas : Hempstead Co. : Hope, 16-VI-1935 (Museum of Comparative Zoology, Cambridge).

    Arkansas : Hempstead Co. : Hope, 19-VI-1931 (Cornell University).

    Arkansas : Hempstead Co. : Hope, VII-1926 (Museum of Comparative Zoology, Cambridge).

    Arkansas : Hempstead Co. : Hope, 31-VII-1930 (California Academy of Science).

    Arkansas : Independence Co. : Newark, 23-VI-1886 (National Museum of Natural History, Washington).

    Arkansas : Merkel, 26-II-1909 (National Museum of Natural History, Washington).

    Arkansas : Johnson Co., 5-VI-1975 (University of Arkansas).

    Arkansas : Johnson Co., 23-VI-1977 (University of Arkansas).

    Arkansas : Lafayette Co., 2/3-VI-1965 (University of Arkansas).

    Arkansas : Lawrence Co. : no further data (California Academy of Science).

    Arkansas : Imboden (California Academy of Science).

    Arkansas : Imboden, 25-V-1929 (California Academy of Science)

    Arkansas : Imboden, 13-VI-1931 (California Academy of Science)

    Arkansas : Imboden, 15-VI-1925 (California Academy of Science)

    Arkansas : Imboden, VII-1926 (California Academy of Science, Field Museum of Natural History)

    Arkansas : Imboden, 13-VII-1925 (Field Museum of Natural History).

    Arkansas : Logan Co. : Magazine Mountain : Cameron Bluff Campground, 28-VI-1990 (American Museum of Natural History, New York).

    Arkansas : Logan Co. : Magazine Mountain : Cameron Bluff Campground, 18-VII-1990 (American Museum of Natural History, New York).

    Arkansas : Mississippi Co. : 6-VI-1971 (University of Arkansas).

    Arkansas : Montgomery Co. : Upper Lake Ouachita (National Museum of Natural History, Washington).

    Arkansas : Montgomery Co. : Upper Lake Ouachita, 6-VIII-1961 (National Museum of Natural History, Washington).

    Arkansas : Phillips Co. : Helena, 16-VI-1920 (Mississippi State University).

    Arkansas : Pulaski Co. : Little Rock, 26-VI-1983 (Florida State Collection of Arthropods).

    Arkansas : Pulaski Co. : Maumalle.

    Arkansas : Savoy, 20-VII-1988 (University of Arkansas)

    Arkansas : Sebastian Co. : Fort Smith, 13-VII-1983, light trap (Florida State Collection of Arthropods).

    Arkansas : Sebastian Co. : Fort Smith, 1-VII-July 1984 (Florida State Collection of Arthropods).

    Arkansas : Sevier Co., VIII-1972 (University of Arkansas).

    Arkansas : St. Francis Co. : Forrest City, VI-1965 (National Museum of Natural History, Washington).

    Arkansas : Washington Co., 1938 (University of Arkansas).

    Arkansas : Washington Co., 3-VI-1977 (University of Arkansas).

    Arkansas : Washington Co., 20-VI-1964 (University of Arkansas).

    Arkansas : Washington Co., 30-VII-1971 (University of Arkansas).

    Arkansas : Washington Co., 8-X-1976 (University of Arkansas).

    Arkansas : Washington Co., 4-XII (University of Arkansas).

    Tennessee (Natural History Museum, London, California Academy of Science).

    Tennessee : Bedford Co. (National Museum of Natural History, Washington).

    Tennessee : Benton Co. (Carnegie Museum of Natural History, Florida State Collection of Arthropods, National Museum of Natural History, Washington).

    Tennessee : Can-oil Co. (Florida State Collection of Arthropods, National Museum of Natural History, Washington).

    Tennessee : Coffee Co. (University of Tennessee).

    Tennessee : Davidson Co. (National Museum of Natural History, Washington).

    Tennessee : Dickson Co. (Florida State Collection of Arthropods).

    Tennessee : Gibson Co. (University of Tennessee).

    Tennessee : Hamilton Co. (Florida State Collection of Arthropods, National Museum of Natural History, Washington).

    Tennessee : Hardeman Co. (CMNM).

    Tennessee : Knox Co. (University of Tennessee, Field Museum of Natural History).

    Tennessee : Marion Co. (Carnegie Museum of Natural History).

    Tennessee : Morgan Co. (California Academy of Science, Carnegie Museum of Natural History, Field Museum of Natural History, Florida State Collection of Arthropods, Museum of Comparative Zoology, Cambridge, Manchester Museum, University of Kentucky, National Museum of Natural History, Washington).

    Tennessee : Obion Co. (Illinois Natural History Survey).

    Tennessee : Roane. Co. (Carnegie Museum of Natural History, Museum of Comparative Zoology, Cambridge).

    Tennessee : Rutherford Co. (University of Michigan).

    Tennessee : Sevier Co. (Texas A & M University).

    Tennessee : Shelby Co. (Carnegie Museum of Natural History).

    North Carolina (California Academy of Science, Field Museum of Natural History, Museum of Comparative Zoology, Cambridge, Florida State Collection of Arthropods, National Museum of Natural History, Washington).

    North Carolina : Bertie Co. (North Carolina State University).

    North Carolina : Cabarrus Co. (California Academy of Science, Florida State Collection of Arthropods, North Carolina State University).

    North Carolina : Camden Co. (North Carolina State University).

    North Carolina : Carteret Co. (Florida State Collection of Arthropods).

    North Carolina : Catawba Co. (North Carolina State University).

    North Carolina : Chatham Co. (North Carolina State University).

    North Carolina : Cleveland Co. (North Carolina State University).

    North Carolina : Craven Co. (Florida State Collection of Arthropods).

    North Carolina : Davidson Co. (North Carolina State University).

    North Carolina : Duplin Co. (North Carolina State University).

    North Carolina : Durham Co. (North Carolina State University).

    North Carolina : Edgecomb Co. (North Carolina State University).

    North Carolina : Franklin Co. (North Carolina State University).

    North Carolina : Gaston Co (Museum of Comparative Zoology, Cambridge, North Carolina State University).

    North Carolina : Graham Co. (California Academy of Science).

    North Carolina : Granville Co. (North Carolina State University, University of Kansas, University of Nebraska].

    North Carolina : Halifax Co. (National Museum of Natural History, Washington).

    North Carolina : Henderson Co. (North Carolina State University).

    North Carolina : Iredell Co. (North Carolina State University).

    North Carolina : Jackson Co. (Ohio State University).

    North Carolina : Johnston Co. (North Carolina State University, Florida State Collection of Arthropods).

    North Carolina : Macon Co. (North Carolina State University).

    North Carolina : Mecklenburg Co. (Natural History Museum, London, North Carolina State University).

    North Carolina : Moore Co. (North Carolina State University, Ohio State University).

    North Carolina : Nash Co. (National Museum of Natural History, Washington).

    North Carolina : Person Co. (North Carolina State University, University of Minnesota).

    North Carolina : Pitt Co. (University of Kansas).

    North Carolina : Randolph Co. (Cornell University).

    North Carolina : Scotland Co. (National Museum of Natural History, Washington).

    North Carolina : Shelby Co., 35°15N-81°35W : State Forest

    North Carolina : Stanly Co. (Texas A & M University).

    North Carolina : Swain Co. (Cornell University).

    North Carolina : Union Co. : Waxhaw, 34°55N-80°45W

    North Carolina : Union Co. (Carnegie Museum of Natural History, North Carolina State University).

    North Carolina : Wake Co. (Field Museum of Natural History, Florida State Collection of Arthropods, North Carolina State University, University of Michigan).

    North Carolina : Washington Co. (Florida State Collection of Arthropods).

    North Carolina : Wayne Co. (North Carolina State University).

     

    South Carolina : Tarplay Gap, 4-VII-1956 (Field Museum of Natural History).

    South Carolina : Anderson Co. : Pendleton, 3-VI-1927 (Manchester Museum).

    South Carolina : Bamburg Co. : IX-1927 (Clemson University).

    South Carolina : Calhoun Co. : Congaree Swamp (Carnegie Museum of Natural History).

    South Carolina : Fail-field Co. : Parr (citated in Kirk, 1970).

    South Carolina : Fail-field Co. : Winnsboro (citated in Kirk, 1970).

    South Carolina : Florence Co. : Florence (citated in Kirk, 1969).

    South Carolina : Greenville Co. : Greer, 3-VII-1931 (Clemson University).

    South Carolina : Lancaster Co. : Richburg, 10-VI-1939 (National Museum of Natural History, Washington).

    South Carolina : Laurens Co. : Enoree (Carnegie Museum of Natural History).

    South Carolina : Lexington Co. : Lexington, 1-VI-1982 (Clemson University).

    South Carolina : McCormick Co. : Willington (citated in Kirk, 1970).

    South Carolina : Oconee Co. : CCC Camp, VII-1943 (National Museum of Natural History, Washington).

    South Carolina : Oconee State Park, 18-III-1974, reared- emerged 6-IX-1974 (Texas A & M University).

    South Carolina : Pickens Co. : Clemson, 6-VI-1975 (Texas A & M University).

    South Carolina : Pickens Co. : Clemson, 15-VI-1975 (

  • fairmairei Planet, 1897

    4 photos

     

    Lucanus fairmairei

     

    Lucanus (Lucanus) fairmairei PLANET, 1897.

    Planet, 1897:265; Planet, 1902:80; Nagel, 1929:117; Arrow, 1950:54, pl.4; Benesh, 1960:141; Kurosawa, 1978:143; Maes, 1992:17; Mizunuma & Nagai, 1994:217, pl.17.

    Myanmar = Burma : Chin State (N. Chin Hills);

    China : Xizang = Thibet* (Se-Pin-Lou-Chan, Ya-Tcheou), Yunnan;

    Thailand : Chiang Mai (Nan).

     

    TEXT : MAES JEAN-MICHEL

     

    PICTURES : GALANT ALAIN

     

  • fanjingshanus Huang & Chen, 2010

    1 photo

     

    Lucanus fanjingshanus

     

     

    Lucanus fanjingshanus , HUANG H. &  CHEN C. , 2010

    Ø       China, Guizhou province

                                                                                                                                                

    Bibliography :

     

    HUANG H. &  CHEN C. , ( 2010 ) Stag Beetles of China I ; Formosa Ecological Compagy, 288 p ; page 123-125 .

     

    TEXT : GALANT ALAIN 

     

    PICTURES : HUANG HAO

     

  • formosanus Planet, 1899

    6 photos

     

    Lucanus (Lucanus) formosanus

     

     

    Lucanus (Lucanus) formosanus PLANET, 1899.

    Planet, 1899:36; Planet, 1902:42; Mollenkamp, 1913:19; Miwa, 1931:321; Benesh, 1960:141; Lacroix, 1972:54; De Lisle, 1973:137-138; Kurosawa, 1978:143; Sakaino, 1981:23; Klausnitzer, 1982:45; Hashimoto, 1990:23-26, pl.4; Imanishi, 1990:15; Maes, 1992:17; Ma, 1993:280; Sakaino, Yu & Chu, 1994:122; Mizunuma & Nagai, 1994:215, pl.13; Kawano, 1997:454.

    Taiwan = Formose* : Taoyuan Distr. (Mt. Rara Shan), Hualien Distr. (Pilu, Hungye near Juisui), Nantou Distr. (Jiyuetan, Puli = Hori), Kaoshiung Distr. (Liukuei), Taitung Distr. (Chihpen wenchuan), ? (Meishi, Suisharyo).

    Note : Ma (1993) cites this species from China (Hunan--Wuling Mts.).

     

    TEXT : MAES JEAN-MICHEL 

     

    PICTURES : GALANT ALAIN

     

  • fortunei Saunders, 1854

    5 photos

     

    Lucanus (Lucanus) fortunei

     

     

    Lucanus (Lucanus) fortunei SAUNDERS, 1854.

    Saunders, 1854:46, pl.3; Planet, 1902:85, pl.15; Heyne & Taschenberg, 1908:50; Nagel, 1926:117

    Arrow, 1943:134; Didier & Séguy, 1952:pl.18; Benesh, 1960:141; Kurosawa, 1978:143; Bomans, 1989:4, 9; Maes, 1992:17; Mizunuma & Nagai, 1994:216, pl.16.

    China* : Jiangsu, Shanghai (Shanghai), Jiangxi, Zhejiang = Tche-Kiang (Hang-Tcheou, Mt. Siming Shan), Fujian = Fokien (Yun-Ling-Shan, San-Chiang, Sanming, Mt. Taiyun Shan, Kuantun), Guangdong, ? (Tche-Ping--Simeon-Ten).

     

    TEXT : MAES JEAN-MICHEL 

     

    PICTURES : GALANT ALAIN

     

  • fryi ssp. fryi Boileau, 1911

    7 photos

     

    Lucanus (Lucanus) fryi

     

     

    Lucanus (Lucanus) fryi BOILEAU, 1911.

    Lucanus fryi BOILEAU, 1911:431, pl.34 [Burma : Ruby Mines]

    Lucanus fryi BOILEAU; Arrow, 1950:48, pl.4-5

    Lucanus fryi BOILEAU; Didier & Séguy, 1952:pl.17

    Lucanus fryi BOILEAU; Benesh, 1960:141

    Lucanus fryi BOILEAU; Lacroix, 1971:561

    Lucanus fryi BOILEAU; Bomans, 1991:146

    Lucanus fryi BOILEAU; Maes, 1992:17

    Lucanus fryi BOILEAU; Kawano, 1994:9

    Lucanus fryi BOILEAU; Mizunuma & Nagai, 1994:213, pl.11

    Lucanus fryi BOILEAU; Kawano, 1997:454.

    Lucanus fryi BOILEAU; Araya, 15, figs.38-39.

    •Myanmar = Burma : Ruby Mines, col. Doherty, ex coll. Fry 1905-100 (male HOLOTYPE of Lucanus fryi BOILEAU, 1911 in Natural History Museum, London) (male PARATYPE of Lucanus fryi BOILEAU, 1911 in Museum National d'Histoire Naturelle, Paris).
    •Myanmar : Kambaiti
    •China : Yunnan : Pe-Yen-Tsing
    •Thailand : Chiang Mai : Fang
    •Thailand : Chiang Mai : Fang : Mt. Doi Pahompok Bibliography :
    ARAYA K. (2001) Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae) from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.

     

    TEXT : MAES JEAN-MICHEL 

     

    PICTURES : GALANT ALAIN

     

  • fujianensis Schenk, 2008

    4 photos

     

    Lucanus ( Lucanus )  fujianensis

    Ø      

     

          Lucanus ( Lucanus ) fujianensis, SCHENK 2008

          

          China, Fujian

                                                                                                                                                

    Bibliography :

     

    SCHENK ( 2008 ) Contribution to the knowledge of the Stag beetles of Asia (Coleoptera, Lucanidae) and description of several new taxa, Lucanus angusticornis inclinatus. Beetles world, 2008; No.1, P1-14, 2008, > ISSN 1867-2892)

     

     

    TEXT : GALANT ALAIN

     

    PICTURES : GALANT ALAIN

  • gamunus Sawada & Watanabe, 1960

    4 photos

     

    Lucanus (Lucanus) gamunus

     

     

    Lucanus (Lucanus) gamunus SAWADA & WATANABE, 1960 [gammunus, gammus].

    Sawada & Watanabe, 1960:99; Nomura, 1960:39; Kurosawa, 1978:141-153; Ishida, 1987:5; Kojima, 1987:43; Isida & Fujioka, 1988:3; Araya, 1989:6; Téllez, 1990:39; Nakata, 1991:22-25; Maes, 1992:17; Mizunuma & Nagai, 1994:217, pl.17; Araya, 1996:20.

    Japan : Tokyo Met. [Izu Is. (Kozujima I., Mikurajima I. {Sato})].

     

    TEXT : MAES JEAN-MICHEL

     

    PICTURES : GALANT ALAIN

     

  • hayashii Nagai, 2000

    4 photos

     

    Lucanus hayashii

     

     

    Lucanus hayashii , NAGAI S. 2003.

    Ø       Myanmar  Katchin Putao Kushin

                                                                                                                                                

    Bibliography :

     

    NAGAI S. ( 2000 ) Twelve New Species, Three New Subspecies, Two New Status and with the Check List of the Family Lucnidae of Northern Myanmar. The Insect of Myanmar, 2000; pp 73-108 Plate 19-30

     

    GEKKAN MUSHI ( 2003 ) Suite a The Lucanid beetles of the world.de MIZUNUMA T. & NAGAI S. Gekkan Mushi, 390: 2-17

     

    HUANG H. ( 2006 ) New descriptions and notes on Chinese stag-beetles, with discovery of the second species of Noseolucanus from SE Tibet. Coleoptera, 10(2006): 11-34, 19 figg.

     

    TEXT : GALANT ALAIN 

     

    PICTURES : GALANT ALAIN

     

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